{"id":35931,"date":"2020-04-08T09:58:25","date_gmt":"2020-04-08T06:58:25","guid":{"rendered":"http:\/\/eranistis.net\/wordpress\/?p=35931"},"modified":"2020-04-16T17:39:56","modified_gmt":"2020-04-16T14:39:56","slug":"%ce%bf%ce%bb%ce%bf%ce%ba%ce%bb%ce%b7%cf%81%cf%89%ce%bc%ce%ad%ce%bd%ce%b7-%ce%b1%ce%bd%cf%84%ce%b9%ce%bc%ce%b5%cf%84%cf%8e%cf%80%ce%b9%cf%83%ce%b7-%cf%84%cf%89%ce%bd-%ce%ba%ce%bf%cf%81%ce%bf%ce%bd","status":"publish","type":"post","link":"https:\/\/eranistis.net\/wordpress\/2020\/04\/08\/%ce%bf%ce%bb%ce%bf%ce%ba%ce%bb%ce%b7%cf%81%cf%89%ce%bc%ce%ad%ce%bd%ce%b7-%ce%b1%ce%bd%cf%84%ce%b9%ce%bc%ce%b5%cf%84%cf%8e%cf%80%ce%b9%cf%83%ce%b7-%cf%84%cf%89%ce%bd-%ce%ba%ce%bf%cf%81%ce%bf%ce%bd\/","title":{"rendered":"\u039f\u03bb\u03bf\u03ba\u03bb\u03b7\u03c1\u03c9\u03bc\u03ad\u03bd\u03b7 \u03b1\u03bd\u03c4\u03b9\u03bc\u03b5\u03c4\u03ce\u03c0\u03b9\u03c3\u03b7 \u03c4\u03c9\u03bd \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03ce\u03bd \u03c4\u03bf\u03bd 21\u03bf \u03b1\u03b9\u03ce\u03bd\u03b1"},"content":{"rendered":"

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\"\u039c\u03ad\u03c7\u03c1\u03b9<\/a>
\u039c\u03ad\u03c7\u03c1\u03b9 \u03c4\u03b1 \u03c4\u03ad\u03bb\u03b7 \u03c4\u03bf\u03c5 2019 \u03b3\u03bd\u03c9\u03c1\u03af\u03b6\u03b1\u03bc\u03b5 6 \u03c0\u03b1\u03b8\u03bf\u03b3\u03cc\u03bd\u03bf\u03c5\u03c2 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03cd\u03c2, \u03bf\u03b9 \u03bf\u03c0\u03bf\u03af\u03bf\u03b9 \u03c0\u03c1\u03bf\u03ba\u03b1\u03bb\u03bf\u03cd\u03c3\u03b1\u03bd \u03b1\u03c3\u03b8\u03ad\u03bd\u03b5\u03b9\u03b5\u03c2 \u03ae \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b5\u03c2. \u0395\u03be \u03b1\u03c5\u03c4\u03ce\u03bd \u03bf\u03b9 4 \u03c0\u03c1\u03bf\u03ba\u03b1\u03bb\u03bf\u03cd\u03bd \u03bc\u03b9\u03b1 \u03ae\u03c0\u03b9\u03b1 \u03c3\u03c5\u03bd\u03ae\u03b8\u03c9\u03c2 \u03bd\u03cc\u03c3\u03bf \u03cc\u03c0\u03c9\u03c2 \u03c4\u03bf \u03ba\u03bf\u03b9\u03bd\u03cc \u03ba\u03c1\u03c5\u03bf\u03bb\u03cc\u03b3\u03b7\u03bc\u03b1 \u03c3\u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf \u03ba\u03b1\u03b9 \u03b5\u03af\u03bd\u03b1\u03b9 \u03b3\u03bd\u03c9\u03c3\u03c4\u03bf\u03af \u03c9\u03c2 \u03bf\u03b9 \u03b9\u03bf\u03af hCoV-229E, hCoV-HKU1, hCoV-NL63 \u03ba\u03b1\u03b9 hCoV-OC43.<\/figcaption><\/figure>\n

O\u03c1\u03b9\u03c3\u03bc\u03ad\u03bd\u03b1 \u03c0\u03b5\u03b9\u03c1\u03ac\u03bc\u03b1\u03c4\u03b1 \u03b3\u03b5\u03bd\u03b5\u03c4\u03b9\u03ba\u03ae\u03c2 \u03bc\u03b7\u03c7\u03b1\u03bd\u03b9\u03ba\u03ae\u03c2 \u03ba\u03b1\u03b9 \u03b3\u03b5\u03bd\u03b5\u03c4\u03b9\u03ba\u03ae\u03c2 \u03bc\u03b5\u03c4\u03b1\u03c4\u03c1\u03bf\u03c0\u03ae\u03c2 \u03b5\u03b3\u03b5\u03af\u03c1\u03bf\u03c5\u03bd \u03b6\u03b7\u03c4\u03ae\u03bc\u03b1\u03c4\u03b1 \u03b2\u03b9\u03bf\u03b1\u03c3\u03c6\u03ac\u03bb\u03b5\u03b9\u03b1\u03c2 \u03ba\u03b1\u03b9 \u03c0\u03b9\u03b8\u03b1\u03bd\u03cc\u03c4\u03b7\u03c4\u03b1\u03c2 \u03b4\u03b9\u03b1\u03c3\u03c0\u03bf\u03c1\u03ac\u03c2 \u03b5\u03b9\u03b4\u03ce\u03bd \u03c0\u03bf\u03c5 \u03b4\u03b7\u03bc\u03b9\u03bf\u03c5\u03c1\u03b3\u03ae\u03b8\u03b7\u03ba\u03b1\u03bd in vitro. \u039c\u03b5\u03c4\u03ac \u03b1\u03c0\u03cc \u03c0\u03bf\u03bb\u03bb\u03ad\u03c2 \u03b4\u03b5\u03ba\u03b1\u03b5\u03c4\u03af\u03b5\u03c2 \u03c3\u03c4\u03b7\u03bd \u0395\u03bb\u03bb\u03ac\u03b4\u03b1 \u03c0\u03b1\u03c1\u03b1\u03c4\u03b7\u03c1\u03ae\u03b8\u03b7\u03ba\u03b1\u03bd \u03b8\u03ac\u03bd\u03b1\u03c4\u03bf\u03b9 \u03b1\u03c0\u03cc \u03be\u03b5\u03c7\u03b1\u03c3\u03bc\u03ad\u03bd\u03b5\u03c2 \u03bd\u03cc\u03c3\u03bf\u03c5\u03c2, (\u03c0.\u03c7. \u03b5\u03bb\u03bf\u03bd\u03bf\u03c3\u03af\u03b1, \u03c6\u03c5\u03bc\u03b1\u03c4\u03af\u03c9\u03c3\u03b7 \u03b1\u03c0\u03cc \u03c4\u03b1 \u03c3\u03c6\u03b1\u03b3\u03b5\u03af\u03b1 \u03ba\u03b1\u03b9 \u03c4\u03b1 \u03b6\u03ce\u03b1, \u03b4\u03b9\u03c6\u03b8\u03b5\u03c1\u03af\u03c4\u03b9\u03b4\u03b1), \u03ba\u03b1\u03b8\u03ce\u03c2 \u03c0\u03bf\u03bb\u03bb\u03ac \u03c0\u03c1\u03bf\u03b3\u03c1\u03ac\u03bc\u03bc\u03b1\u03c4\u03b1 \u03ba\u03b1\u03c4\u03b1\u03c0\u03bf\u03bb\u03ad\u03bc\u03b7\u03c3\u03b7\u03c2 \u03ae \u03b5\u03c0\u03b1\u03b3\u03c1\u03cd\u03c0\u03bd\u03b7\u03c3\u03b7\u03c2 \u03c0\u03b1\u03c1\u03b1\u03c3\u03af\u03c4\u03c9\u03bd \u03ba\u03b1\u03b9 \u03b6\u03c9\u03bf\u03bd\u03cc\u03c3\u03c9\u03bd \u03c5\u03c0\u03bf\u03b2\u03b1\u03b8\u03bc\u03af\u03c3\u03c4\u03b7\u03ba\u03b1\u03bd \u03ae \u03b5\u03b3\u03ba\u03b1\u03c4\u03b1\u03bb\u03b5\u03af\u03c6\u03b8\u03b7\u03ba\u03b1\u03bd \u03c3\u03c4\u03b1\u03b4\u03b9\u03b1\u03ba\u03ac \u03bc\u03b5\u03c4\u03ac \u03c4\u03bf 1960 \u03b3\u03b9\u03b1 \u03bb\u03cc\u03b3\u03bf\u03c5\u03c2 \u03bf\u03b9\u03ba\u03bf\u03bd\u03bf\u03bc\u03b9\u03ba\u03bf\u03cd\u03c2 \u03ba\u03b1\u03b9 \u03ac\u03bb\u03bb\u03bf\u03c5\u03c2. \u0397 \u03b4\u03b7\u03bc\u03cc\u03c3\u03b9\u03b1 \u03c5\u03b3\u03b5\u03af\u03b1 \u03c0\u03ad\u03c1\u03b1\u03c3\u03b5 \u03c3\u03b5 \u03b4\u03b5\u03cd\u03c4\u03b5\u03c1\u03bf \u03c0\u03bb\u03ac\u03bd\u03bf, \u03cc\u03c0\u03c9\u03c2 \u03ba\u03b1\u03b9 \u03b7 \u03b2\u03b1\u03c3\u03b9\u03ba\u03ae \u03bc\u03b5\u03bb\u03ad\u03c4\u03b7 (\u03b6\u03c9\u03bf\u03b3\u03b5\u03c9\u03b3\u03c1\u03b1\u03c6\u03af\u03b1, \u03c3\u03c5\u03b3\u03ba\u03c1\u03b9\u03c4\u03b9\u03ba\u03ae \u03c6\u03c5\u03c3\u03b9\u03bf\u03bb\u03bf\u03b3\u03af\u03b1 \u03c4\u03c9\u03bd \u03b5\u03b9\u03b4\u03ce\u03bd, \u03c3\u03c5\u03c3\u03c4\u03b7\u03bc\u03b1\u03c4\u03b9\u03ba\u03ae \u03c4\u03c9\u03bd \u03b5\u03bd\u03b4\u03b7\u03bc\u03b9\u03ba\u03ce\u03bd \u03b5\u03b9\u03b4\u03ce\u03bd \u03ba\u03ac\u03b8\u03b5 \u03c0\u03b5\u03c1\u03b9\u03bf\u03c7\u03ae\u03c2, \u03c0\u03b1\u03c1\u03b1\u03c3\u03b9\u03c4\u03bf\u03bb\u03bf\u03b3\u03af\u03b1).<\/p>\n

\u039f\u03b9 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03af \u03ba\u03b1\u03b9 \u03b9\u03b4\u03af\u03c9\u03c2 \u03c4\u03bf\u03c5 \u03b3\u03ad\u03bd\u03bf\u03c5\u03c2 betacoronovirus \u03ad\u03c7\u03bf\u03c5\u03bd \u03bc\u03b5\u03bb\u03b5\u03c4\u03b7\u03b8\u03b5\u03af \u03bc\u03b5\u03c4\u03ac \u03c4\u03bf 1960. \u0388\u03c7\u03bf\u03c5\u03bd \u03c3\u03b1\u03bd \u03ba\u03bf\u03b9\u03bd\u03cc \u03c4\u03bf\u03c5\u03c2 \u03c0\u03c1\u03cc\u03b3\u03bf\u03bd\u03bf \u03ad\u03bd\u03b1\u03bd \u03b9\u03cc \u03c0\u03bf\u03c5 \u03b1\u03bd\u03b9\u03c7\u03bd\u03b5\u03cd\u03b8\u03b7\u03ba\u03b5 \u03c3\u03c4\u03b9\u03c2 \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03b5\u03c2 \u03ba\u03b1\u03b9 \u03bc\u03b5\u03c4\u03b1\u03c6\u03ad\u03c1\u03b8\u03b7\u03c3\u03b1\u03bd \u03c3\u03b5 \u03c0\u03bf\u03bb\u03bb\u03ac \u03ac\u03bb\u03bb\u03b1 \u03b5\u03af\u03b4\u03b7 \u03b8\u03b7\u03bb\u03b1\u03c3\u03c4\u03b9\u03ba\u03ce\u03bd.<\/strong><\/p>\n

\u039f 21\u03bf\u03c2 <\/sup>\u03b1\u03b9\u03ce\u03bd\u03b1\u03c2 \u03be\u03b5\u03ba\u03af\u03bd\u03b7\u03c3\u03b5 \u03bc\u03b5 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b5\u03c2 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03ce\u03bd (CoVs). \u03a4\u03bf \u03bf\u03be\u03cd \u03b1\u03bd\u03b1\u03c0\u03bd\u03b5\u03c5\u03c3\u03c4\u03b9\u03ba\u03cc \u03c3\u03cd\u03bd\u03b4\u03c1\u03bf\u03bc\u03bf \u03c0\u03c1\u03bf\u03ad\u03ba\u03c5\u03c8\u03b5 \u03b1\u03c0\u03cc \u03c4\u03bf\u03bd \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03cc SARS-CoV \u03c4\u03bf 2002 \u03ba\u03b1\u03b9 2003, \u03ba\u03b1\u03b9 \u03c4\u03bf \u03b1\u03bd\u03b1\u03c0\u03bd\u03b5\u03c5\u03c3\u03c4\u03b9\u03ba\u03cc \u03c3\u03cd\u03bd\u03b4\u03c1\u03bf\u03bc\u03bf \u03c4\u03b7\u03c2 \u039c\u03ad\u03c3\u03b7\u03c2 \u0391\u03bd\u03b1\u03c4\u03bf\u03bb\u03ae\u03c2 \u03c0\u03c1\u03bf\u03ad\u03ba\u03c5\u03c8\u03b5 \u03b1\u03c0\u03cc \u03c4\u03bf\u03bd \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03cc MERS-CoV \u03c4\u03bf 2012 \u03ba\u03b1\u03b9 2015. \u03a3\u03c4\u03bf \u03b7\u03bb\u03b5\u03ba\u03c4\u03c1\u03bf\u03bd\u03b9\u03ba\u03cc \u03bc\u03b9\u03ba\u03c1\u03bf\u03c3\u03ba\u03cc\u03c0\u03b9\u03bf \u03bf\u03b9 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03af \u03c6\u03b1\u03af\u03bd\u03b5\u03c4\u03b1\u03b9 \u03bd\u03b1 \u03c6\u03ad\u03c1\u03bf\u03c5\u03bd \u03ad\u03bd\u03b1 \u03c3\u03c4\u03ad\u03bc\u03bc\u03b1 (corona) \u03b3\u03cd\u03c1\u03c9 \u03c4\u03bf\u03c5\u03c2, \u03bb\u03cc\u03b3\u03c9 \u03c4\u03b7\u03c2 \u03c0\u03b1\u03c1\u03bf\u03c5\u03c3\u03af\u03b1\u03c2 \u03c0\u03ac\u03bd\u03c9 \u03b1\u03c0\u03cc \u03c4\u03bf\u03bd \u03c6\u03ac\u03ba\u03b5\u03bb\u03bf\/\u03c0\u03b5\u03c1\u03af\u03b2\u03bb\u03b7\u03bc\u03bc\u03b1 \u03c4\u03bf\u03c5 \u03b9\u03bf\u03cd \u03c0\u03bf\u03bb\u03bb\u03ce\u03bd \u03c3\u03b5\u03b9\u03c1\u03ce\u03bd \u03b1\u03c0\u03cc \u03b1\u03ba\u03c4\u03b9\u03bd\u03c9\u03c4\u03ad\u03c2 \u03b3\u03bb\u03c5\u03ba\u03bf\u03c0\u03c1\u03c9\u03c4\u03b5\u03ca\u03bd\u03b5\u03c2 (spikes) \u03bc\u03b5 \u03c4\u03b9\u03c2 \u03bf\u03c0\u03bf\u03af\u03b5\u03c2 \u03c0\u03c1\u03bf\u03c3\u03ba\u03bf\u03bb\u03bb\u03ce\u03bd\u03c4\u03b1\u03b9 \u03c3\u03c4\u03bf \u03ba\u03cd\u03c4\u03c4\u03b1\u03c1\u03bf \u03c4\u03bf\u03c5 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ae \u03b3\u03b9\u03b1 \u03bd\u03b1 \u03b5\u03b9\u03c3\u03ad\u03bb\u03b8\u03bf\u03c5\u03bd \u03c3\u03b5 \u03b1\u03c5\u03c4\u03cc, \u03bc\u03b5\u03c4\u03ac \u03c4\u03b7\u03bd \u03bc\u03cc\u03bb\u03c5\u03bd\u03c3\u03b7. \u039c\u03b5\u03c4\u03b1\u03bb\u03bb\u03ac\u03be\u03b5\u03b9\u03c2 \u03c3\u03c4\u03b9\u03c2 \u03c0\u03c1\u03c9\u03c4\u03b5\u03ca\u03bd\u03b5\u03c2 \u03b1\u03c5\u03c4\u03ad\u03c2, \u03b1\u03bb\u03bb\u03ac\u03b6\u03bf\u03c5\u03bd \u03ba\u03b1\u03b9 \u03c4\u03b7\u03bd \u03bc\u03bf\u03bb\u03c5\u03c3\u03bc\u03b1\u03c4\u03b9\u03ba\u03cc\u03c4\u03b7\u03c4\u03b1 \u03c4\u03bf\u03c5 \u03b9\u03bf\u03cd.<\/p>\n

\u039f\u03b9 \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03b5\u03c2 \u03b1\u03bd\u03ae\u03ba\u03bf\u03c5\u03bd \u03c3\u03c4\u03b7\u03bd \u03c4\u03ac\u03be\u03b7 \u03c4\u03c9\u03bd Chiropter\u03b1, \u03b5\u03af\u03bd\u03b1\u03b9 \u03bd\u03c5\u03c7\u03c4\u03cc\u03b2\u03b9\u03b1, \u03ba\u03bf\u03c3\u03bc\u03bf\u03c0\u03bf\u03bb\u03af\u03c4\u03b9\u03ba\u03b1 \u03b8\u03b7\u03bb\u03b1\u03c3\u03c4\u03b9\u03ba\u03ac \u03c0\u03bf\u03c5 \u03c0\u03b5\u03c4\u03bf\u03cd\u03bd. \u03a0\u03bf\u03bb\u03bb\u03ac \u03b5\u03af\u03b4\u03b7 \u03c4\u03bf\u03c5\u03c2 \u03bc\u03b5\u03c4\u03b1\u03bd\u03b1\u03c3\u03c4\u03b5\u03cd\u03bf\u03c5\u03bd \u03ba\u03b1\u03b9 \u03ac\u03bb\u03bb\u03b1 \u03c0\u03ad\u03c6\u03c4\u03bf\u03c5\u03bd \u03c3\u03b5 \u03c7\u03b5\u03b9\u03bc\u03ad\u03c1\u03b9\u03b1 \u03bd\u03ac\u03c1\u03ba\u03b7. \u03a0\u03c1\u03cc\u03ba\u03b5\u03b9\u03c4\u03b1\u03b9 \u03b3\u03b9\u03b1 \u03bc\u03b9\u03b1 \u03c4\u03ac\u03be\u03b7 \u03c0\u03bf\u03bb\u03c5\u03c0\u03bb\u03b7\u03b8\u03ae \u03bc\u03b5 1.200 \u03b5\u03af\u03b4\u03b7. \u03a3\u03cd\u03bc\u03c6\u03c9\u03bd\u03b1 \u03bc\u03b5 \u03c4\u03bf\u03c5\u03c2 Hu et al. (2015) \u03bf\u03b9 \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03b5\u03c2 \u03ad\u03c7\u03bf\u03c5\u03bd \u03bc\u03b9\u03b1 \u03bc\u03b5\u03b3\u03ac\u03bb\u03b7 \u03b3\u03b5\u03c9\u03b3\u03c1\u03b1\u03c6\u03b9\u03ba\u03ae \u03ba\u03b1\u03c4\u03b1\u03bd\u03bf\u03bc\u03ae \u03ba\u03b1\u03b9 \u03b5\u03af\u03bd\u03b1\u03b9 \u03c6\u03c5\u03c3\u03b9\u03ba\u03bf\u03af \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ad\u03c2 \u03c0\u03bf\u03bb\u03bb\u03ce\u03bd \u03ba\u03b1\u03b9 \u03b4\u03b9\u03b1\u03c6\u03bf\u03c1\u03b5\u03c4\u03b9\u03ba\u03ce\u03bd \u03b9\u03ce\u03bd \u03bc\u03b5 \u03bc\u03b5\u03b3\u03ac\u03bb\u03b7 \u03c5\u03b3\u03b5\u03b9\u03bf\u03bd\u03bf\u03bc\u03b9\u03ba\u03ae \u03c3\u03b7\u03bc\u03b1\u03c3\u03af\u03b1 \u03b3\u03b9\u03b1 \u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf. \u0395\u03bd\u03b4\u03b5\u03b9\u03ba\u03c4\u03b9\u03ba\u03ac \u03b1\u03bd\u03b1\u03c6\u03ad\u03c1\u03bf\u03bd\u03c4\u03b1\u03b9 \u03c9\u03c2 \u03c6\u03c5\u03c3\u03b9\u03ba\u03bf\u03af \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ad\u03c2 \u03c4\u03c9\u03bd \u03b9\u03ce\u03bd (lyssaviruses, paramyxoviruses, coronaviruses, filoviruses) \u03c0\u03bf\u03c5 \u03c0\u03c1\u03bf\u03ba\u03b1\u03bb\u03bf\u03cd\u03bd \u03c4\u03b7 \u03bb\u03cd\u03c3\u03c3\u03b1, \u03b9\u03bb\u03b1\u03c1\u03ac, \u03c0\u03b1\u03c1\u03c9\u03c4\u03af\u03c4\u03b9\u03b4\u03b1, \u03ba.\u03b1. \u0397 \u03bb\u03cd\u03c3\u03c3\u03b1 \u03c0\u03b1\u03c1\u03b1\u03bc\u03ad\u03bd\u03b5\u03b9 \u03bc\u03b9\u03b1 \u03c3\u03bf\u03b2\u03b1\u03c1\u03ae \u03bd\u03cc\u03c3\u03bf\u03c2 \u03b1\u03c0\u03cc \u03c4\u03b7\u03bd \u03bf\u03c0\u03bf\u03af\u03b1 \u03c0\u03b5\u03b8\u03b1\u03af\u03bd\u03bf\u03c5\u03bd \u03b4\u03b5\u03ba\u03ac\u03b4\u03b5\u03c2 \u03c7\u03b9\u03bb\u03b9\u03ac\u03b4\u03b5\u03c2 \u03ac\u03c4\u03bf\u03bc\u03b1 \u03b5\u03c4\u03b7\u03c3\u03af\u03c9\u03c2. \u03a4\u03bf 2015 \u03c0\u03ad\u03b8\u03b1\u03bd\u03b1\u03bd 17.400 \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf\u03b9, 40% \u03b1\u03c5\u03c4\u03ce\u03bd \u03c0\u03b1\u03b9\u03b4\u03b9\u03ac, \u03ba\u03c5\u03c1\u03af\u03c9\u03c2 \u03c3\u03c4\u03b7\u03bd \u0391\u03c3\u03af\u03b1 \u03ba\u03b1\u03b9 \u0391\u03c6\u03c1\u03b9\u03ba\u03ae.<\/p>\n

H \u03b7\u03b8\u03bf\u03bb\u03bf\u03b3\u03af\u03b1 \u03c4\u03c9\u03bd \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03c9\u03bd (\u03b6\u03ce\u03b1 \u03ba\u03bf\u03b9\u03bd\u03c9\u03bd\u03b9\u03ba\u03ac \u03ba\u03b1\u03b9 \u03bc\u03b1\u03ba\u03c1\u03cc\u03b2\u03b9\u03b1, \u03bc\u03b5\u03c4\u03b1\u03ba\u03b9\u03bd\u03bf\u03cd\u03bc\u03b5\u03bd\u03b1 \u03b3\u03c1\u03ae\u03b3\u03bf\u03c1\u03b1) \u03c4\u03b9\u03c2 \u03ba\u03ac\u03bd\u03b5\u03b9 \u03c0\u03bf\u03bb\u03cd \u03b1\u03c0\u03bf\u03c4\u03b5\u03bb\u03b5\u03c3\u03bc\u03b1\u03c4\u03b9\u03ba\u03ad\u03c2 \u03c3\u03c4\u03b7\u03bd \u03b4\u03b9\u03b1\u03c3\u03c0\u03bf\u03c1\u03ac \u03bc\u03bf\u03bb\u03c5\u03c3\u03bc\u03b1\u03c4\u03b9\u03ba\u03ce\u03bd \u03b1\u03c3\u03b8\u03b5\u03bd\u03b5\u03b9\u03ce\u03bd. \u0398\u03b7\u03c1\u03b5\u03c5\u03c4\u03ad\u03c2 \u03c4\u03bf\u03c5\u03c2 \u03b5\u03af\u03bd\u03b1\u03b9 \u03c4\u03bf \u03b3\u03b5\u03c1\u03ac\u03ba\u03b9, \u03b7 \u03ba\u03bf\u03c5\u03ba\u03bf\u03c5\u03b2\u03ac\u03b3\u03b9\u03b1, \u03c4\u03bf \u03ba\u03bf\u03c5\u03bd\u03ac\u03b2\u03b9, \u03b7 \u03b3\u03ac\u03c4\u03b1 \u03ba\u03b1\u03b9 \u03c4\u03b1 \u03c6\u03af\u03b4\u03b9\u03b1. \u0391\u03bd\u03ac\u03bc\u03b5\u03c3\u03b1 \u03c3\u03c4\u03bf\u03c5\u03c2 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03cd\u03c2 Alphacoronavirus \u03ba\u03b1\u03b9 Betacoronavirus \u03c0\u03bf\u03c5 \u03c0\u03c1\u03bf\u03c3\u03b2\u03ac\u03bb\u03bb\u03bf\u03c5\u03bd \u03b8\u03b7\u03bb\u03b1\u03c3\u03c4\u03b9\u03ba\u03ac, \u03bf\u03b9 7 \u03c3\u03c4\u03bf\u03c5\u03c2 15 \u03ad\u03c7\u03bf\u03c5\u03bd \u03b2\u03c1\u03b5\u03b8\u03b5\u03af \u03c3\u03c4\u03b9\u03c2 \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03b5\u03c2, \u03bf\u03b9 \u03bf\u03c0\u03bf\u03af\u03b5\u03c2 \u03b5\u03af\u03bd\u03b1\u03b9 \u03bf\u03b9 \u03ba\u03cd\u03c1\u03b9\u03bf\u03b9 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ad\u03c2 \u03c4\u03bf\u03c5\u03c2 \u03ba\u03b1\u03b9 \u03bf\u03b9 \u03bf\u03c0\u03bf\u03af\u03b5\u03c2 \u03c6\u03b9\u03bb\u03bf\u03be\u03b5\u03bd\u03bf\u03cd\u03bd \u03c0\u03bf\u03bb\u03bb\u03bf\u03cd\u03c2 \u03c0\u03b5\u03c1\u03b9\u03c3\u03c3\u03cc\u03c4\u03b5\u03c1\u03bf\u03c5\u03c2 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03cd\u03c2 \u03b1\u03c0\u03cc \u03ba\u03ac\u03b8\u03b5 \u03ac\u03bb\u03bb\u03bf \u03b5\u03af\u03b4\u03bf\u03c2, \u03b5\u03bd\u03ce \u03c6\u03b1\u03af\u03bd\u03b5\u03c4\u03b1\u03b9 \u03bd\u03b1 \u03c0\u03b1\u03af\u03b6\u03bf\u03c5\u03bd \u03ad\u03bd\u03b1 \u03c3\u03b7\u03bc\u03b1\u03bd\u03c4\u03b9\u03ba\u03cc \u03c1\u03cc\u03bb\u03bf \u03c3\u03c4\u03b7\u03bd \u03b3\u03b5\u03bd\u03b5\u03c4\u03b9\u03ba\u03ae \u03c4\u03bf\u03c5\u03c2 \u03b5\u03be\u03ad\u03bb\u03b9\u03be\u03b7 (Hu et al., 2015).<\/strong><\/p>\n

\u039f\u03b9 \u03b6\u03c9\u03bf\u03bd\u03cc\u03c3\u03bf\u03b9 \u03b5\u03af\u03bd\u03b1\u03b9 \u03bd\u03cc\u03c3\u03bf\u03b9 \u03c0\u03bf\u03c5 \u03bc\u03b5\u03c4\u03b1\u03b4\u03af\u03b4\u03bf\u03bd\u03c4\u03b1\u03b9 \u03c3\u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf \u03b1\u03c0\u03cc \u03c4\u03b1 \u03b6\u03ce\u03b1, \u03c4\u03bf\u03c5\u03c2 \u03b5\u03bd\u03b4\u03b9\u03ac\u03bc\u03b5\u03c3\u03bf\u03c5\u03c2 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ad\u03c2 \u03c4\u03bf\u03c5\u03c2. \u03a4\u03b1 \u03b6\u03ce\u03b1 \u03b1\u03c0\u03bf\u03c4\u03b5\u03bb\u03bf\u03cd\u03bd \u03b1\u03c0\u03bf\u03b8\u03ae\u03ba\u03b5\u03c2 \u03c4\u03bf\u03c5 \u03b9\u03bf\u03cd \u03c3\u03c4\u03b7 \u03c6\u03cd\u03c3\u03b7 \u03b3\u03b9\u03b1 \u03c4\u03b7\u03bd \u03b4\u03b9\u03ac\u03b4\u03bf\u03c3\u03ae \u03c4\u03bf\u03c5 \u03b1\u03bb\u03bb\u03ac \u03ba\u03b1\u03b9 \u03c4\u03b7\u03bd \u03b5\u03be\u03ad\u03bb\u03b9\u03be\u03ae \u03c4\u03bf\u03c5 \u03bc\u03ad\u03c3\u03c9 \u03bc\u03b5\u03c4\u03b1\u03bb\u03bb\u03ac\u03be\u03b5\u03c9\u03bd \u03c3\u03b5 \u03bd\u03ad\u03b1 \u03c3\u03c4\u03b5\u03bb\u03ad\u03c7\u03b7. \u039c\u03ad\u03c7\u03c1\u03b9 \u03c4\u03b1 \u03c4\u03ad\u03bb\u03b7 \u03c4\u03bf\u03c5 2019 \u03b3\u03bd\u03c9\u03c1\u03af\u03b6\u03b1\u03bc\u03b5 6 \u03c0\u03b1\u03b8\u03bf\u03b3\u03cc\u03bd\u03bf\u03c5\u03c2 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03cd\u03c2, \u03bf\u03b9 \u03bf\u03c0\u03bf\u03af\u03bf\u03b9 \u03c0\u03c1\u03bf\u03ba\u03b1\u03bb\u03bf\u03cd\u03c3\u03b1\u03bd \u03b1\u03c3\u03b8\u03ad\u03bd\u03b5\u03b9\u03b5\u03c2 \u03ae \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b5\u03c2. \u0395\u03be \u03b1\u03c5\u03c4\u03ce\u03bd \u03bf\u03b9 4 \u03c0\u03c1\u03bf\u03ba\u03b1\u03bb\u03bf\u03cd\u03bd \u03bc\u03b9\u03b1 \u03ae\u03c0\u03b9\u03b1 \u03c3\u03c5\u03bd\u03ae\u03b8\u03c9\u03c2 \u03bd\u03cc\u03c3\u03bf \u03cc\u03c0\u03c9\u03c2 \u03c4\u03bf \u03ba\u03bf\u03b9\u03bd\u03cc \u03ba\u03c1\u03c5\u03bf\u03bb\u03cc\u03b3\u03b7\u03bc\u03b1 \u03c3\u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf \u03ba\u03b1\u03b9 \u03b5\u03af\u03bd\u03b1\u03b9 \u03b3\u03bd\u03c9\u03c3\u03c4\u03bf\u03af \u03c9\u03c2 \u03bf\u03b9 \u03b9\u03bf\u03af hCoV-229E, hCoV-HKU1, hCoV-NL63 \u03ba\u03b1\u03b9 hCoV-OC43. \u0384\u039f\u03bb\u03bf\u03b9 \u03b1\u03c5\u03c4\u03bf\u03af \u03ad\u03c7\u03bf\u03c5\u03bd \u03bc\u03b9\u03ba\u03c1\u03ae \u03c3\u03b7\u03bc\u03b1\u03c3\u03af\u03b1 \u03b3\u03b9\u03b1 \u03c4\u03b7\u03bd \u03b4\u03b7\u03bc\u03cc\u03c3\u03b9\u03b1 \u03c5\u03b3\u03b5\u03af\u03b1 (Weston & Frieman, 2020).<\/p>\n

\u039f\u03b9 \u03ac\u03bb\u03bb\u03bf\u03b9 \u03b4\u03cd\u03bf \u03cc\u03bc\u03c9\u03c2 (SARS-CoV, MERS-CoV ) \u03c0\u03c1\u03bf\u03ba\u03b1\u03bb\u03bf\u03cd\u03bd \u03c4\u03bf \u03c0\u03b1\u03b3\u03ba\u03cc\u03c3\u03bc\u03b9\u03bf \u03b5\u03bd\u03b4\u03b9\u03b1\u03c6\u03ad\u03c1\u03bf\u03bd, \u03c4\u03cc\u03c3\u03bf \u03b3\u03b9\u03b1 \u03c4\u03b7\u03bd \u03b2\u03b1\u03c1\u03cd\u03c4\u03b7\u03c4\u03b1 \u03c4\u03b7\u03c2 \u03bd\u03cc\u03c3\u03bf\u03c5 \u03c0\u03bf\u03c5 \u03c0\u03c1\u03bf\u03ba\u03b1\u03bb\u03bf\u03cd\u03bd, \u03c4\u03bf\u03bd \u03b6\u03c9\u03bf\u03bd\u03bf\u03c4\u03b9\u03ba\u03cc \u03c4\u03bf\u03c5\u03c2 \u03ba\u03cd\u03ba\u03bb\u03bf, \u03cc\u03c3\u03bf \u03ba\u03b1\u03b9 \u03b3\u03b9\u03b1 \u03c4\u03b7\u03bd \u03c0\u03b1\u03b3\u03ba\u03cc\u03c3\u03bc\u03b9\u03b1 \u03bf\u03b9\u03ba\u03bf\u03bd\u03bf\u03bc\u03af\u03b1 (Huai-Dong et al., 2005, Lau et al., 2005).<\/strong><\/p>\n

\"\u0391\u03bd\u03ac\u03bc\u03b5\u03c3\u03b1<\/a>
\u0391\u03bd\u03ac\u03bc\u03b5\u03c3\u03b1 \u03c3\u03c4\u03bf\u03c5\u03c2 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03cd\u03c2 Alphacoronavirus \u03ba\u03b1\u03b9 Betacoronavirus \u03c0\u03bf\u03c5 \u03c0\u03c1\u03bf\u03c3\u03b2\u03ac\u03bb\u03bb\u03bf\u03c5\u03bd \u03b8\u03b7\u03bb\u03b1\u03c3\u03c4\u03b9\u03ba\u03ac, \u03bf\u03b9 7 \u03c3\u03c4\u03bf\u03c5\u03c2 15 \u03ad\u03c7\u03bf\u03c5\u03bd \u03b2\u03c1\u03b5\u03b8\u03b5\u03af \u03c3\u03c4\u03b9\u03c2 \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03b5\u03c2, \u03bf\u03b9 \u03bf\u03c0\u03bf\u03af\u03b5\u03c2 \u03b5\u03af\u03bd\u03b1\u03b9 \u03bf\u03b9 \u03ba\u03cd\u03c1\u03b9\u03bf\u03b9 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ad\u03c2 \u03c4\u03bf\u03c5\u03c2 \u03ba\u03b1\u03b9 \u03bf\u03b9 \u03bf\u03c0\u03bf\u03af\u03b5\u03c2 \u03c6\u03b9\u03bb\u03bf\u03be\u03b5\u03bd\u03bf\u03cd\u03bd \u03c0\u03bf\u03bb\u03bb\u03bf\u03cd\u03c2 \u03c0\u03b5\u03c1\u03b9\u03c3\u03c3\u03cc\u03c4\u03b5\u03c1\u03bf\u03c5\u03c2 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03cd\u03c2 \u03b1\u03c0\u03cc \u03ba\u03ac\u03b8\u03b5 \u03ac\u03bb\u03bb\u03bf \u03b5\u03af\u03b4\u03bf\u03c2, \u03b5\u03bd\u03ce \u03c6\u03b1\u03af\u03bd\u03b5\u03c4\u03b1\u03b9 \u03bd\u03b1 \u03c0\u03b1\u03af\u03b6\u03bf\u03c5\u03bd \u03ad\u03bd\u03b1 \u03c3\u03b7\u03bc\u03b1\u03bd\u03c4\u03b9\u03ba\u03cc \u03c1\u03cc\u03bb\u03bf \u03c3\u03c4\u03b7\u03bd \u03b3\u03b5\u03bd\u03b5\u03c4\u03b9\u03ba\u03ae \u03c4\u03bf\u03c5\u03c2 \u03b5\u03be\u03ad\u03bb\u03b9\u03be\u03b7 (Hu et al., 2015).<\/figcaption><\/figure>\n

\u03a4\u03bf \u03bf\u03be\u03cd \u03b1\u03bd\u03b1\u03c0\u03bd\u03b5\u03c5\u03c4\u03b9\u03ba\u03cc \u03c3\u03cd\u03bd\u03b4\u03c1\u03bf\u03bc\u03bf \u03c0\u03bf\u03c5 \u03c0\u03c1\u03bf\u03ba\u03bb\u03ae\u03b8\u03b7\u03ba\u03b5 \u03c4\u03bf 2002 \u03b1\u03c0\u03cc \u03c4\u03bf\u03bd SARS-CoV, \u03c0\u03c1\u03bf\u03ba\u03ac\u03bb\u03b5\u03c3\u03b5 \u03c4\u03bf \u03c0\u03b1\u03b3\u03ba\u03cc\u03c3\u03bc\u03b9\u03bf \u03b5\u03bd\u03b4\u03b9\u03b1\u03c6\u03ad\u03c1\u03bf\u03bd \u03b1\u03c6\u03bf\u03cd \u03c3\u03b5 \u03bb\u03af\u03b3\u03bf\u03c5\u03c2 \u03bc\u03ae\u03bd\u03b5\u03c2 \u03b1\u03c5\u03c4\u03cc\u03c2 \u03bf \u03b9\u03cc\u03c2 \u03c4\u03c9\u03bd \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03c9\u03bd \u03c0\u03ad\u03c1\u03b1\u03c3\u03b5 \u03c3\u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf. \u0397 \u03bc\u03b5\u03c4\u03b1\u03c6\u03bf\u03c1\u03ac \u03b1\u03c5\u03c4\u03ae \u03ad\u03b3\u03b9\u03bd\u03b5 \u03bc\u03b5 \u03b5\u03bd\u03b4\u03b9\u03ac\u03bc\u03b5\u03c3\u03bf \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ae (Drexler et al., 2014) \u03bc\u03b5 \u03c4\u03bf \u03cc\u03bd\u03bf\u03bc\u03b1 \u03bc\u03bf\u03c3\u03c7\u03bf\u03b3\u03b1\u03bb\u03ae (\u03ad\u03bd\u03b1 \u03b8\u03b7\u03bb\u03b1\u03c3\u03c4\u03b9\u03ba\u03cc \u03c0\u03bf\u03c5 \u03bc\u03bf\u03b9\u03ac\u03b6\u03b5\u03b9 \u03bc\u03b5 \u03b3\u03ac\u03c4\u03b1 \u03ae \u03ba\u03bf\u03c5\u03bd\u03ac\u03b2\u03b9, \u03b6\u03b5\u03af \u03c3\u03c4\u03b7\u03bd \u0391\u03c3\u03af\u03b1 \u03ba\u03b1\u03b9 \u03c4\u03b7\u03bd \u0391\u03c6\u03c1\u03b9\u03ba\u03ae \u03ba\u03b1\u03b9 \u03b8\u03c1\u03ad\u03c6\u03b5\u03c4\u03b1\u03b9 \u03bc\u03b5 \u03c6\u03c1\u03bf\u03cd\u03c4\u03b1 \u03ba\u03b1\u03b9 \u03ad\u03bd\u03c4\u03bf\u03bc\u03b1, \u03ba\u03b1\u03b9 \u03b1\u03c0\u03bf\u03c4\u03b5\u03bb\u03b5\u03af \u03c4\u03c1\u03bf\u03c6\u03ae \u03ba\u03b1\u03b9 \u03ba\u03b1\u03c4\u03bf\u03b9\u03ba\u03af\u03b4\u03b9\u03bf \u03b3\u03b9\u03b1 \u03c4\u03bf\u03c5\u03c2 \u0391\u03c3\u03b9\u03ac\u03c4\u03b5\u03c2). \u0397 \u03bc\u03b5\u03c4\u03ac\u03b4\u03bf\u03c3\u03b7 \u03b1\u03c0\u03cc \u03c4\u03b7\u03bd \u03bc\u03bf\u03c3\u03c7\u03bf\u03b3\u03b1\u03bb\u03ae \u03c3\u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf \u03ad\u03b3\u03b9\u03bd\u03b5 \u03c3\u03c4\u03b7\u03bd Guangdong \u03c4\u03b7\u03c2 \u039a\u03af\u03bd\u03b1\u03c2. O SARS-CoV \u03b1\u03c0\u03bf\u03bc\u03bf\u03bd\u03ce\u03b8\u03b7\u03ba\u03b5 \u03b1\u03c0\u03cc \u03c4\u03b7\u03bd \u03bc\u03bf\u03c3\u03c7\u03bf\u03b3\u03b1\u03bb\u03ae Himalayan palm civets (Paguma<\/em> larvata<\/em>) \u03ba\u03b1\u03b9 \u03b1\u03c0\u03cc \u03c4\u03bf raccoon dog (Nyctereutes<\/em> procyonoides<\/em>) \u03c3\u03c4\u03b7\u03bd \u03b1\u03b3\u03bf\u03c1\u03ac Shenzhen \u03c4\u03bf 2002\u20132003. \u039f\u03b9 \u03b1\u03bb\u03bb\u03b7\u03bb\u03bf\u03c5\u03c7\u03af\u03b5\u03c2 \u03c4\u03bf\u03c5\u03c2 \u03ad\u03b4\u03c9\u03c3\u03b1\u03bd \u03bf\u03bc\u03bf\u03b9\u03cc\u03c4\u03b7\u03c4\u03b1 99.8% identity \u03bc\u03b5 \u03c4\u03bf\u03bd SARS-CoV \u03b1\u03c0\u03cc \u03b1\u03bd\u03b8\u03c1\u03ce\u03c0\u03bf\u03c5\u03c2 (Huai-Dong et al., 2005). \u039f \u03b9\u03cc\u03c2 \u03b1\u03c5\u03c4\u03cc\u03c2 \u03bc\u03cc\u03bb\u03c5\u03bd\u03b5 8.000 \u03ac\u03c4\u03bf\u03bc\u03b1 \u03bc\u03b5 \u03b8\u03bd\u03b7\u03c3\u03b9\u03bc\u03cc\u03c4\u03b7\u03c4\u03b1 10% \u03c0\u03b5\u03c1\u03af\u03c0\u03bf\u03c5, \u03b5\u03bd\u03ce \u03c4\u03bf \u03b5\u03c0\u03cc\u03bc\u03b5\u03bd\u03bf \u03ad\u03c4\u03bf\u03c2 2003 \u03b7 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b1 \u03b1\u03c5\u03c4\u03ae \u03ad\u03c3\u03b2\u03b7\u03c3\u03b5.<\/p>\n

To 2012 \u03ad\u03bd\u03b1 \u03bd\u03ad\u03bf \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03b9\u03ba\u03cc \u03bf\u03be\u03cd \u03b1\u03bd\u03b1\u03c0\u03bd\u03b5\u03c5\u03c3\u03c4\u03b9\u03ba\u03cc \u03c3\u03cd\u03bd\u03b4\u03c1\u03bf\u03bc\u03bf \u03b1\u03c0\u03cc \u03c4\u03bf\u03bd MERS-CoV \u03b5\u03bc\u03c6\u03b1\u03bd\u03af\u03c3\u03c4\u03b7\u03ba\u03b5 \u03c3\u03c4\u03b7\u03bd \u03a3. \u0391\u03c1\u03b1\u03b2\u03af\u03b1. \u03a4\u03b1 \u03bb\u03af\u03b3\u03b1 \u03c0\u03b5\u03c1\u03b9\u03c3\u03c4\u03b1\u03c4\u03b9\u03ba\u03ac \u03c4\u03b7\u03c2 \u03bd\u03cc\u03c3\u03bf\u03c5 \u03c3\u03c4\u03b7\u03bd \u0391\u03c3\u03af\u03b1, \u0395\u03c5\u03c1\u03ce\u03c0\u03b7 \u03ba\u03b1\u03b9 \u0397\u03a0\u0391 \u03c3\u03c5\u03c3\u03c7\u03b5\u03c4\u03af\u03c3\u03c4\u03b7\u03ba\u03b1\u03bd \u03bc\u03b5 \u03ac\u03c4\u03bf\u03bc\u03b1 \u03c0\u03bf\u03c5 \u03b5\u03af\u03c7\u03b1\u03bd \u03c4\u03b1\u03be\u03b9\u03b4\u03ad\u03c8\u03b5\u03b9 \u03c3\u03c4\u03b7\u03bd \u03a3. \u0391\u03c1\u03b1\u03b2\u03af\u03b1. \u0397 \u03b4\u03b5\u03cd\u03c4\u03b5\u03c1\u03b7 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b1 \u03c4\u03b7\u03c2 \u03bd\u03cc\u03c3\u03bf\u03c5 \u03c3\u03c4\u03b7\u03bd \u039a\u03bf\u03c1\u03ad\u03b1 \u03c4\u03bf 2015 \u03bc\u03b5 185 \u03c0\u03b5\u03c1\u03b9\u03c3\u03c4\u03b1\u03c4\u03b9\u03ba\u03ac, \u03c0\u03ac\u03bb\u03b9 \u03c3\u03c5\u03bd\u03b4\u03ad\u03b8\u03b7\u03ba\u03b5 \u03bc\u03b5 \u03c4\u03b1\u03be\u03b9\u03b4\u03b9\u03ce\u03c4\u03b7 \u03b1\u03c0\u03cc \u03c4\u03b7\u03bd \u039c\u03ad\u03c3\u03b7 \u0391\u03bd\u03b1\u03c4\u03bf\u03bb\u03ae.<\/strong> \u0397 \u03bc\u03b5\u03c4\u03ac\u03b4\u03bf\u03c3\u03b7 \u03c4\u03bf\u03c5 MERS-CoV \u03b1\u03c0\u03cc \u03c4\u03b9\u03c2 \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03b5\u03c2 \u03c3\u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf, \u03c3\u03b5 \u03b1\u03c5\u03c4\u03ae\u03bd \u03c4\u03b7\u03bd \u03c0\u03b5\u03c1\u03af\u03c0\u03c4\u03c9\u03c3\u03b7 \u03ad\u03b3\u03b9\u03bd\u03b5 \u03bc\u03ad\u03c3\u03c9 \u03b5\u03bd\u03cc\u03c2 \u03ac\u03bb\u03bb\u03bf\u03c5 \u03b5\u03bd\u03b4\u03b9\u03ac\u03bc\u03b5\u03c3\u03bf\u03c5 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ae, \u03c4\u03b7\u03bd \u03ba\u03b1\u03bc\u03ae\u03bb\u03b1. \u039c\u03b5\u03bb\u03ad\u03c4\u03b5\u03c2 \u03ad\u03b4\u03b5\u03b9\u03be\u03b1\u03bd \u03cc\u03c4\u03b9 \u03bf MERS-CoV \u03bc\u03c0\u03bf\u03c1\u03b5\u03af \u03bd\u03b1 \u03bc\u03bf\u03bb\u03cd\u03bd\u03b5\u03b9 \u03ba\u03b1\u03b9 \u03bd\u03b1 \u00ab\u03c0\u03bf\u03bb\u03bb\u03b1\u03c0\u03bb\u03b1\u03c3\u03b9\u03b1\u03c3\u03c4\u03b5\u03af\u00bb \u03c3\u03b5 \u03ba\u03cd\u03c4\u03c4\u03b1\u03c1\u03b1 \u03b1\u03c0\u03cc \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf, \u03ac\u03bb\u03bb\u03b1 \u03c0\u03c1\u03c9\u03c4\u03b5\u03cd\u03bf\u03bd\u03c4\u03b1, \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03b5\u03c2, \u03c7\u03bf\u03af\u03c1\u03bf\u03c5\u03c2, \u03b1\u03af\u03b3\u03b5\u03c2, \u03ac\u03bb\u03bf\u03b3\u03b1, \u03ba\u03b1\u03bc\u03ae\u03bb\u03b5\u03c2, \u03ba\u03bf\u03c5\u03bd\u03ad\u03bb\u03b9\u03b1 \u03ba.\u03b1. (Hu et al., 2015). \u0391\u03be\u03b9\u03bf\u03c3\u03b7\u03bc\u03b5\u03af\u03c9\u03c4\u03bf \u03b5\u03af\u03bd\u03b1\u03b9 \u03cc\u03c4\u03b9 \u03b1\u03c5\u03c4\u03ae \u03b7 \u03b4\u03b9\u03b1\u03b5\u03b9\u03b4\u03b9\u03ba\u03ae \u03bc\u03b5\u03c4\u03b1\u03c0\u03ae\u03b4\u03b7\u03c3\u03b7 \u03c4\u03bf\u03c5 \u03b9\u03bf\u03cd \u03ad\u03b3\u03b9\u03bd\u03b5 \u03b4\u03b5\u03ba\u03b1\u03b5\u03c4\u03af\u03b5\u03c2 \u03c0\u03c1\u03b9\u03bd \u03c4\u03b7\u03bd \u03b5\u03bc\u03c6\u03ac\u03bd\u03b9\u03c3\u03b7 \u03c4\u03b7\u03c2 \u03bd\u03cc\u03c3\u03bf\u03c5. \u039f \u03b9\u03cc\u03c2 MERS-CoV \u03b1\u03bd\u03b9\u03c7\u03bd\u03b5\u03cd\u03b8\u03b7\u03ba\u03b5 \u03c3\u03b5 \u03ba\u03b1\u03bc\u03ae\u03bb\u03b5\u03c2 \u03b1\u03c0\u03cc \u03c4\u03b7\u03bd \u03a3. \u0391\u03c1\u03b1\u03b2\u03af\u03b1, \u03c4\u03bf \u039a\u03b1\u03c4\u03ac\u03c1, \u03c4\u03b7\u03bd \u0391\u03af\u03b3\u03c5\u03c0\u03c4\u03bf \u03ba\u03b1\u03b9 \u03ad\u03c7\u03b5\u03b9 \u03bf\u03bc\u03bf\u03b9\u03cc\u03c4\u03b7\u03c4\u03b1 >99%, \u03bc\u03b5 \u03c4\u03b9\u03c2 \u03b1\u03bb\u03bb\u03b7\u03bb\u03bf\u03c5\u03c7\u03af\u03b5\u03c2 \u03b1\u03c0\u03cc \u03c4\u03bf\u03bd MERS-CoV \u03b1\u03c0\u03cc \u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf. \u0391\u03c0\u03cc \u03c6\u03c5\u03bb\u03bf\u03b3\u03b5\u03bd\u03b5\u03c4\u03b9\u03ba\u03ad\u03c2 \u03bc\u03b5\u03bb\u03ad\u03c4\u03b5\u03c2 \u03c6\u03b1\u03af\u03bd\u03b5\u03c4\u03b1\u03b9 \u03c0\u03b9\u03b8\u03b1\u03bd\u03cc\u03bd \u03cc\u03c4\u03b9 \u03bf \u03c0\u03b9\u03bf \u03ba\u03bf\u03bd\u03c4\u03b9\u03bd\u03cc\u03c2 \u03c0\u03c1\u03cc\u03b3\u03bf\u03bd\u03bf\u03c2 \u03c4\u03bf\u03c5 \u03b9\u03bf\u03cd \u03bc\u03b5\u03c4\u03b1\u03c0\u03ae\u03b4\u03b7\u03c3\u03b5 \u03b1\u03c0\u03cc \u03c4\u03b9\u03c2 \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03b5\u03c2 \u03c3\u03c4\u03b9\u03c2 \u03ba\u03b1\u03bc\u03ae\u03bb\u03b5\u03c2 \u03c3\u03c4\u03b7\u03bd \u0391\u03c6\u03c1\u03b9\u03ba\u03ae 20 \u03c7\u03c1\u03cc\u03bd\u03b9\u03b1 \u03c0\u03c1\u03b9\u03bd, \u03cc\u03c4\u03b1\u03bd \u03bc\u03b5\u03c4\u03b1\u03c6\u03ad\u03c1\u03b8\u03b7\u03ba\u03b1\u03bd \u03ba\u03b1\u03bc\u03ae\u03bb\u03b5\u03c2 \u03b1\u03c0\u03cc \u03c4\u03b7\u03bd \u0391\u03c1\u03b1\u03b2\u03b9\u03ba\u03ae \u03c7\u03b5\u03c1\u03c3\u03cc\u03bd\u03b7\u03c3\u03bf. \u039f \u03b9\u03cc\u03c2 \u03b1\u03c5\u03c4\u03cc\u03c2 \u03c7\u03b1\u03c1\u03b1\u03ba\u03c4\u03b7\u03c1\u03af\u03b6\u03b5\u03c4\u03b1\u03b9 \u03b1\u03c0\u03cc \u03bc\u03b9\u03ba\u03c1\u03cc\u03c4\u03b5\u03c1\u03b7 \u03bc\u03b5\u03c4\u03b1\u03b4\u03bf\u03c4\u03b9\u03ba\u03cc\u03c4\u03b7\u03c4\u03b1 \u03b1\u03c0\u03cc \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf \u03c3\u03b5 \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf, \u03b1\u03ba\u03cc\u03bc\u03b1 \u03ba\u03b1\u03b9 \u03b1\u03c0\u03cc \u03c4\u03bf SARS-CoV, \u03bc\u03cc\u03bb\u03c5\u03bd\u03b5 \u03bc\u03cc\u03bb\u03b9\u03c2 2.500 \u03ac\u03c4\u03bf\u03bc\u03b1 \u03ba\u03c5\u03c1\u03af\u03c9\u03c2 \u03c3\u03c4\u03b7\u03bd \u039c\u03ad\u03c3\u03b7 \u0391\u03bd\u03b1\u03c4\u03bf\u03bb\u03ae \u03b1\u03bb\u03bb\u03ac \u03b5\u03af\u03c7\u03b5 \u03c0\u03bf\u03bb\u03cd \u03c5\u03c8\u03b7\u03bb\u03ae \u03b8\u03bd\u03b7\u03c3\u03b9\u03bc\u03cc\u03c4\u03b7\u03c4\u03b1 (\u03c0\u03b5\u03c1\u03af\u03c0\u03bf\u03c5 35%), \u03b3\u03b5\u03b3\u03bf\u03bd\u03cc\u03c2 \u03c0\u03bf\u03c5 \u03ba\u03b1\u03c4\u03b1\u03c4\u03ac\u03c3\u03c3\u03b5\u03b9 \u03c4\u03bf\u03bd MERS-CoV \u03c9\u03c2 \u03ad\u03bd\u03b1\u03bd \u03b1\u03c0\u03cc \u03c4\u03bf\u03c5\u03c2 \u03c0\u03b9\u03bf \u03b8\u03b1\u03bd\u03b1\u03c4\u03b7\u03c6\u03cc\u03c1\u03bf\u03c5\u03c2 \u03b9\u03bf\u03cd\u03c2 \u03c3\u03c4\u03bf\u03bd \u03ba\u03cc\u03c3\u03bc\u03bf. \u0397 \u03b1\u03bd\u03c4\u03b9\u03bc\u03b5\u03c4\u03ce\u03c0\u03b9\u03c3\u03b7 \u03c4\u03bf\u03c5 \u03ad\u03b3\u03b9\u03bd\u03b5 \u03b5\u03c6\u03b9\u03ba\u03c4\u03ae \u03c7\u03c9\u03c1\u03af\u03c2 \u03c0\u03bf\u03bb\u03bb\u03ac \u03bc\u03ad\u03c4\u03c1\u03b1, \u03bb\u03cc\u03b3\u03c9 \u03c4\u03bf\u03c5 \u03b5\u03bd\u03c4\u03bf\u03c0\u03b9\u03c3\u03bc\u03bf\u03cd \u03c4\u03bf\u03c5 \u03ba\u03cd\u03c1\u03b9\u03bf\u03c5 \u03b5\u03bd\u03b4\u03b9\u03ac\u03bc\u03b5\u03c3\u03bf\u03c5 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ae (\u03c4\u03b7\u03c2 \u03ba\u03b1\u03bc\u03ae\u03bb\u03b1\u03c2). \u0397 \u03b1\u03bd\u03b1\u03ba\u03ac\u03bb\u03c5\u03c8\u03b7 \u03b5\u03bc\u03b2\u03bf\u03bb\u03af\u03bf\u03c5 \u03b3\u03b9\u03b1 \u03c4\u03bf\u03c5\u03c2 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03cd\u03c2 \u03b4\u03b5\u03bd \u03ad\u03b3\u03b9\u03bd\u03b5 \u03b1\u03ba\u03cc\u03bc\u03b1 \u03b5\u03c6\u03b9\u03ba\u03c4\u03ae, \u03b1\u03bb\u03bb\u03ac \u03c5\u03c0\u03ae\u03c1\u03be\u03b1\u03bd \u03c0\u03bf\u03bb\u03bb\u03ad\u03c2 \u03c5\u03c0\u03bf\u03c3\u03c7\u03cc\u03bc\u03b5\u03bd\u03b5\u03c2 \u03bc\u03b5\u03bb\u03ad\u03c4\u03b5\u03c2 \u03b3\u03b9\u03b1 \u03b5\u03bc\u03b2\u03cc\u03bb\u03b9\u03bf \u03ad\u03bd\u03b1\u03bd\u03c4\u03b9 \u03c4\u03bf\u03c5 MERS-CoV \u03c3\u03c4\u03b9\u03c2 \u03ba\u03b1\u03bc\u03ae\u03bb\u03b5\u03c2.<\/p>\n

\u0388\u03c4\u03c3\u03b9, \u03c0\u03c1\u03b9\u03bd \u03c4\u03b7\u03bd \u03b5\u03ba\u03b4\u03ae\u03bb\u03c9\u03c3\u03b7 \u03c4\u03b7\u03c2 \u03c3\u03b7\u03bc\u03b5\u03c1\u03b9\u03bd\u03ae\u03c2 \u03c0\u03b1\u03bd\u03b4\u03b7\u03bc\u03af\u03b1\u03c2, \u03ad\u03c7\u03bf\u03c5\u03bc\u03b5 \u03b4\u03cd\u03bf \u03c0\u03b1\u03c1\u03cc\u03bc\u03bf\u03b9\u03bf\u03c5\u03c2 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03cd\u03c2 \u03bd\u03b1 \u03c0\u03c1\u03bf\u03ba\u03b1\u03bb\u03bf\u03cd\u03bd \u03c3\u03bf\u03b2\u03b1\u03c1\u03ad\u03c2 \u03b6\u03c9\u03bf\u03bd\u03bf\u03c4\u03b9\u03ba\u03ad\u03c2 \u03bd\u03cc\u03c3\u03bf\u03c5\u03c2, \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b5\u03c2 \u03bc\u03b5 \u03bf\u03be\u03ad\u03b1 \u03b1\u03bd\u03b1\u03c0\u03bd\u03b5\u03c5\u03c3\u03c4\u03b9\u03ba\u03ac \u03c3\u03cd\u03bd\u03b4\u03c1\u03bf\u03bc\u03b1 \u03ba\u03b1\u03b9 \u03c0\u03c1\u03bf\u03c2 \u03c4\u03bf \u03c0\u03b1\u03c1\u03cc\u03bd \u03b3\u03bd\u03c9\u03c3\u03c4\u03bf\u03cd\u03c2 \u03b5\u03bd\u03b4\u03b9\u03ac\u03bc\u03b5\u03c3\u03bf\u03c5\u03c2 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ad\u03c2: \u03b4\u03cd\u03bf \u03b8\u03b7\u03bb\u03b1\u03c3\u03c4\u03b9\u03ba\u03ac, \u03c4\u03b7\u03bd \u03bc\u03bf\u03c3\u03c7\u03bf\u03b3\u03b1\u03bb\u03ae (Paguma larvata) \u03ba\u03b1\u03b9 \u03c4\u03b7\u03bd \u03b4\u03c1\u03bf\u03bc\u03ac\u03b4\u03b1 (camel).<\/strong><\/p>\n

\u03a3\u03c4\u03b1 \u03c4\u03ad\u03bb\u03b7 \u03c4\u03bf\u03c5 2019 \u03b5\u03bc\u03c6\u03b1\u03bd\u03af\u03c3\u03c4\u03b7\u03ba\u03b5 \u03bf \u03c4\u03c1\u03af\u03c4\u03bf\u03c2 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03cc\u03c2 (SARS-CoV-2), \u03b1\u03c0\u03cc \u03c4\u03b9\u03c2 \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03b5\u03c2 \u03c0\u03ac\u03bb\u03b9, \u03b1\u03bb\u03bb\u03ac \u03bc\u03b5 \u03ac\u03b3\u03bd\u03c9\u03c3\u03c4\u03bf \u03b5\u03bd\u03b4\u03b9\u03ac\u03bc\u03b5\u03c3\u03bf \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ae \u03c0\u03c1\u03bf\u03c2 \u03c4\u03bf \u03c0\u03b1\u03c1\u03cc\u03bd. \u03a5\u03c0\u03ac\u03c1\u03c7\u03bf\u03c5\u03bd \u03c3\u03bf\u03b2\u03b1\u03c1\u03ad\u03c2 \u03b5\u03bd\u03b4\u03b5\u03af\u03be\u03b5\u03b9\u03c2 \u03cc\u03c4\u03b9 \u03c3\u03c5\u03bd\u03b4\u03ad\u03b5\u03c4\u03b1\u03b9 \u03bc\u03b5 \u03c4\u03b7\u03bd \u03b4\u03b7\u03bc\u03cc\u03c3\u03b9\u03b1 \u03b1\u03b3\u03bf\u03c1\u03ac \u03b8\u03b1\u03bb\u03b1\u03c3\u03c3\u03b9\u03bd\u03ce\u03bd \u03c3\u03c4\u03b7\u03bd \u039a\u03af\u03bd\u03b1 \u03ba\u03b1\u03b9 \u03bc\u03ad\u03c3\u03c9 \u03b1\u03c5\u03c4\u03ae\u03c2 \u03ad\u03b3\u03b9\u03bd\u03b5 \u03b7 \u03b4\u03b9\u03b1\u03c3\u03c0\u03bf\u03c1\u03ac \u03c4\u03bf\u03c5 \u03c3\u03c4\u03b7\u03bd \u0393\u03b9\u03bf\u03c5\u03c7\u03ac\u03bd \u03b1\u03c1\u03c7\u03b9\u03ba\u03ac. \u03a4\u03b1 \u03b6\u03ce\u03b1 \u03c0\u03bf\u03c5 \u03c0\u03c9\u03bb\u03bf\u03cd\u03bd\u03c4\u03b1\u03b9 \u03c3\u03c4\u03b9\u03c2 \u03b1\u03b3\u03bf\u03c1\u03ad\u03c2 \u03b8\u03b1\u03bb\u03b1\u03c3\u03c3\u03b9\u03bd\u03ce\u03bd \u03c3\u03c4\u03b7\u03bd \u039a\u03af\u03bd\u03b1 \u03bc\u03c0\u03bf\u03c1\u03b5\u03af \u03bd\u03b1 \u03b5\u03af\u03bd\u03b1\u03b9 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ad\u03c2 \u03c4\u03bf\u03c5. \u039f \u03bc\u03c5\u03c1\u03bc\u03b7\u03b3\u03ba\u03bf\u03c6\u03ac\u03b3\u03bf\u03c2 (\u03b5\u03bd\u03c4\u03bf\u03bc\u03bf\u03c6\u03ac\u03b3\u03bf \u03b8\u03b7\u03bb\u03b1\u03c3\u03c4\u03b9\u03ba\u03cc \u03c4\u03b7\u03c2 \u03c4\u03ac\u03be\u03b7\u03c2 Pholid\u03bfta) \u03b5\u03bc\u03c6\u03b1\u03bd\u03af\u03c3\u03c4\u03b7\u03ba\u03b5 \u03c9\u03c2 \u03bf \u03b5\u03bd\u03b4\u03b9\u03ac\u03bc\u03b5\u03c3\u03bf\u03c2 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ae\u03c2 \u03c4\u03bf\u03c5 \u03bd\u03ad\u03bf\u03c5 \u03b9\u03bf\u03cd SARS-CoV-2, \u03b1\u03bb\u03bb\u03ac \u03b1\u03c5\u03c4\u03cc \u03bc\u03ad\u03bd\u03b5\u03b9 \u03bd\u03b1 \u03b5\u03c0\u03b9\u03b2\u03b5\u03b2\u03b1\u03b9\u03c9\u03b8\u03b5\u03af.<\/p>\n

\u0397 \u03bd\u03ad\u03b1 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b1 \u03b5\u03af\u03c7\u03b5 \u03c0\u03c1\u03bf\u03b2\u03bb\u03b5\u03c6\u03b8\u03b5\u03af. \u039f Menachery \u03ba\u03b1\u03b9 \u03bf\u03b9 \u03c3\u03c5\u03bd\u03b5\u03c1\u03b3\u03ac\u03c4\u03b5\u03c2 \u03c4\u03bf\u03c5, \u03c0.\u03c7. \u03b1\u03c0\u03cc \u03c4\u03bf 2016, \u03bc\u03b5 \u03c0\u03b5\u03b9\u03c1\u03ac\u03bc\u03b1\u03c4\u03b1 \u03b3\u03b5\u03bd\u03b5\u03c4\u03b9\u03ba\u03ae\u03c2 \u03bc\u03b7\u03c7\u03b1\u03bd\u03b9\u03ba\u03ae\u03c2 \u03b5\u03af\u03c7\u03b1\u03bd \u03c0\u03c1\u03bf\u03b2\u03bb\u03ad\u03c8\u03b5\u03b9 \u03cc\u03c4\u03b9 \u03ba\u03ac\u03c0\u03bf\u03b9\u03bf\u03b9 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03af \u03c4\u03c9\u03bd \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03c9\u03bd \u03b1\u03c0\u03cc \u03c4\u03b7\u03bd \u0393\u03b9\u03bf\u03c5\u03c7\u03ac\u03bd \u03bc\u03c0\u03bf\u03c1\u03b5\u03af \u03bd\u03b1 \u03c0\u03c1\u03bf\u03ba\u03b1\u03bb\u03ad\u03c3\u03bf\u03c5\u03bd \u03bd\u03ad\u03b5\u03c2 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b5\u03c2 \u03ba\u03b1\u03b9 \u03bc\u03b5\u03b3\u03ac\u03bb\u03b5\u03c2 \u03ba\u03c1\u03af\u03c3\u03b5\u03b9\u03c2. \u0395\u03bd \u03bf\u03bb\u03af\u03b3\u03bf\u03b9\u03c2, \u03b7 \u03bf\u03bc\u03ac\u03b4\u03b1 \u03c4\u03bf\u03c5 Menachery \u03c3\u03c5\u03bd\u03ad\u03b8\u03b5\u03c3\u03b5 \u03ad\u03bd\u03b1\u03bd \u03c0\u03bb\u03ae\u03c1\u03b7 DNA \u03ba\u03bb\u03ce\u03bd\u03bf \u03bc\u03b5 \u03b2\u03ac\u03c3\u03b7 \u03c4\u03b7\u03bd \u03b1\u03bb\u03bb\u03b7\u03bb\u03bf\u03c5\u03c7\u03af\u03b1 \u03c4\u03bf\u03c5 \u03b9\u03bf\u03cd, \u03c0\u03c1\u03bf\u03ba\u03b5\u03b9\u03bc\u03ad\u03bd\u03bf\u03c5 \u03bd\u03b1 \u03c0\u03c1\u03bf\u03b2\u03bb\u03ad\u03c8\u03b5\u03b9 \u03b1\u03bd \u03ad\u03bd\u03b1\u03c2 SARS \u03c4\u03cd\u03c0\u03bf\u03c5 CoV (\u03c0\u03bf\u03c5 \u03bf\u03bd\u03bf\u03bc\u03ac\u03c3\u03c4\u03b7\u03ba\u03b5 WIV1-CoV \u03ba\u03b1\u03b9 \u03b2\u03c1\u03ad\u03b8\u03b7\u03ba\u03b5 \u03c3\u03b5 \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03b5\u03c2 \u03c3\u03c4\u03b7\u03bd Yunnan) \u03b5\u03af\u03c7\u03b5 \u03c4\u03b7\u03bd \u03c0\u03b9\u03b8\u03b1\u03bd\u03cc\u03c4\u03b7\u03c4\u03b1 \u03bd\u03b1 \u03c0\u03c1\u03bf\u03c3\u03b2\u03ac\u03bb\u03bb\u03b5\u03b9 \u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf.<\/strong> \u039f \u03ba\u03bb\u03ce\u03bd\u03bf\u03c2 \u03b1\u03c0\u03bf\u03b4\u03b5\u03af\u03c7\u03c4\u03b7\u03ba\u03b5 \u03b9\u03ba\u03b1\u03bd\u03cc\u03c2 \u03bd\u03b1 \u03b4\u03b7\u03bc\u03b9\u03bf\u03c5\u03c1\u03b3\u03ae\u03c3\u03b5\u03b9 \u03bc\u03bf\u03bb\u03c5\u03c3\u03bc\u03b1\u03c4\u03b9\u03ba\u03cc \u03b9\u03cc \u03b3\u03b9\u03b1 \u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf \u03c3\u03b5 \u03ba\u03b1\u03bb\u03bb\u03b9\u03ad\u03c1\u03b3\u03b5\u03b9\u03b5\u03c2 \u03ba\u03c5\u03c4\u03c4\u03ac\u03c1\u03c9\u03bd \u03ba\u03b1\u03b9 in vivo.<\/p>\n

\u038c\u03c4\u03b1\u03bd \u03c7\u03c1\u03b7\u03c3\u03b9\u03bc\u03bf\u03c0\u03bf\u03b9\u03ae\u03b8\u03b7\u03ba\u03b5 \u03c4\u03bf \u03b3\u03bf\u03bd\u03af\u03b4\u03b9\u03bf spike \u03c4\u03bf\u03c5 WIV1-CoV \u03c3\u03b5 \u03b4\u03b9\u03b1\u03b3\u03bf\u03bd\u03b9\u03b4\u03b9\u03b1\u03ba\u03ac \u03c0\u03bf\u03bd\u03c4\u03af\u03ba\u03b9\u03b1 \u03c4\u03b1 \u03bf\u03c0\u03bf\u03af\u03b1 \u03b5\u03af\u03c7\u03b1\u03bd, \u03bc\u03b5\u03c4\u03ac \u03b1\u03c0\u03cc \u03b3\u03b5\u03bd\u03b5\u03c4\u03b9\u03ba\u03ae \u03bc\u03b5\u03c4\u03b1\u03c4\u03c1\u03bf\u03c0\u03ae, \u03c4\u03b7 \u03b4\u03c5\u03bd\u03b1\u03c4\u03cc\u03c4\u03b7\u03c4\u03b1 \u03bd\u03b1 \u03b5\u03ba\u03c6\u03c1\u03ac\u03b6\u03bf\u03c5\u03bd \u03c4\u03bf\u03bd \u03c5\u03c0\u03bf\u03b4\u03bf\u03c7\u03ad\u03b1 ACE2 \u03cc\u03c0\u03c9\u03c2 \u03c4\u03b1 \u03b1\u03bd\u03b8\u03c1\u03ce\u03c0\u03b9\u03bd\u03b1 \u03ba\u03cd\u03c4\u03c4\u03b1\u03c1\u03b1, \u03c4\u03b1 \u03c0\u03b5\u03b9\u03c1\u03ac\u03bc\u03b1\u03c4\u03b1 \u03ad\u03b4\u03b5\u03b9\u03be\u03b1\u03bd \u03cc\u03c4\u03b9 \u03bf WIV1-CoV \u03ad\u03c7\u03b5\u03b9 \u03c0\u03c1\u03ac\u03b3\u03bc\u03b1\u03c4\u03b9 \u03bc\u03b5\u03b3\u03ac\u03bb\u03b7 \u03c0\u03b9\u03b8\u03b1\u03bd\u03cc\u03c4\u03b7\u03c4\u03b1 \u03b5\u03bc\u03c6\u03ac\u03bd\u03b9\u03c3\u03b7\u03c2 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03b9\u03ce\u03bd \u03c3\u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf \u03ba\u03b1\u03b9 \u03c4\u03b7\u03bd \u03bc\u03b5\u03c4\u03ac\u03b4\u03bf\u03c3\u03b7 \u03b1\u03bd\u03ac\u03bc\u03b5\u03c3\u03b1 \u03c3\u03b5 \u03b1\u03bd\u03b8\u03c1\u03ce\u03c0\u03bf\u03c5\u03c2. \u0395\u03c0\u03b9\u03c0\u03bb\u03ad\u03bf\u03bd, \u03bc\u03b5\u03bb\u03b5\u03c4\u03ae\u03b8\u03b7\u03ba\u03b1\u03bd \u03c4\u03c1\u03cc\u03c0\u03bf\u03c5\u03c2 \u03bc\u03b5\u03c4\u03c1\u03b9\u03b1\u03c3\u03bc\u03bf\u03cd \u03bc\u03b9\u03b1\u03c2 \u03c4\u03ad\u03c4\u03bf\u03b9\u03b1\u03c2 \u03ba\u03c1\u03af\u03c3\u03b7\u03c2 \u03b1\u03c0\u03cc WIV1-\u03c4\u03cd\u03c0\u03bf\u03c5 \u03b9\u03bf\u03cd\u03c2.<\/p>\n

\u03a0\u03b5\u03b9\u03c1\u03b1\u03bc\u03b1\u03c4\u03b9\u03b6\u03cc\u03bc\u03b5\u03bd\u03b7 \u03bc\u03b5 \u03b3\u03b5\u03bd\u03b5\u03c4\u03b9\u03ba\u03ac \u03c4\u03c1\u03bf\u03c0\u03bf\u03c0\u03bf\u03b9\u03b7\u03bc\u03ad\u03bd\u03b1 \u03c0\u03b5\u03b9\u03c1\u03b1\u03bc\u03b1\u03c4\u03cc\u03b6\u03c9\u03b1 \u03ba\u03b1\u03b9 \u03b5\u03c3\u03c4\u03b9\u03ac\u03b6\u03bf\u03bd\u03c4\u03b1\u03c2 \u03c3\u03c4\u03b7\u03bd \u03b5\u03bd\u03b6\u03c5\u03bc\u03b9\u03ba\u03ae \u03c0\u03c1\u03c9\u03c4\u03b5\u03af\u03bd\u03b7 ACE2 \u03bc\u03b5 \u03b1\u03c6\u03bf\u03c1\u03bc\u03ae \u03c4\u03b9\u03c2 \u03c0\u03c1\u03cc\u03c3\u03c6\u03b1\u03c4\u03b5\u03c2 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b5\u03c2 \u03c4\u03bf\u03c5 \u03ad\u03bc\u03c0\u03bf\u03bb\u03b1, \u03c4\u03b7\u03c2 \u03b3\u03c1\u03af\u03c0\u03c0\u03b7\u03c2 \u03ba\u03b1\u03b9 \u03c4\u03bf\u03c5 MERS-CoV, \u03b7 \u03bf\u03bc\u03ac\u03b4\u03b1 Menachery \u03b8\u03b5\u03c9\u03c1\u03bf\u03cd\u03c3\u03b5 \u03cc\u03c4\u03b9 \u03c5\u03c0\u03bf\u03b5\u03ba\u03c4\u03b9\u03bc\u03ac\u03c4\u03b1\u03b9 \u03b7 \u03ba\u03c1\u03af\u03c3\u03b7 \u03c0\u03bf\u03c5 \u03bc\u03c0\u03bf\u03c1\u03b5\u03af \u03bd\u03b1 \u03b4\u03b7\u03bc\u03b9\u03bf\u03c5\u03c1\u03b3\u03ae\u03c3\u03b5\u03b9 \u03ad\u03bd\u03b1\u03c2 \u03bd\u03ad\u03bf\u03c2 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03cc\u03c2. \u0394\u03b5\u03b4\u03bf\u03bc\u03ad\u03bd\u03b7\u03c2 \u03c4\u03b7\u03c2 \u03b1\u03cd\u03be\u03b7\u03c3\u03b7\u03c2 \u03c4\u03c9\u03bd \u03c4\u03b1\u03be\u03b9\u03b4\u03af\u03c9\u03bd \u03b1\u03bd\u03b1\u03c8\u03c5\u03c7\u03ae\u03c2 \u03ba\u03b1\u03b9 \u03c4\u03c9\u03bd \u03ac\u03bd\u03b9\u03c3\u03c9\u03bd \u03b4\u03c5\u03bd\u03b1\u03c4\u03bf\u03c4\u03ae\u03c4\u03c9\u03bd \u03c4\u03c9\u03bd \u03b9\u03b1\u03c4\u03c1\u03b9\u03ba\u03ce\u03bd \u03c5\u03c0\u03bf\u03b4\u03bf\u03bc\u03ce\u03bd \u03c3\u03b5 \u03b4\u03b9\u03ac\u03c6\u03bf\u03c1\u03b5\u03c2 \u03c7\u03ce\u03c1\u03b5\u03c2, \u03c0\u03c1\u03cc\u03c4\u03b5\u03b9\u03bd\u03b5 \u03c4\u03b7\u03bd \u03b1\u03bd\u03ac\u03c0\u03c4\u03c5\u03be\u03b7 \u03bc\u03b5\u03bb\u03b5\u03c4\u03ce\u03bd \u03b3\u03b9\u03b1 \u03c4\u03b7\u03bd \u03bc\u03b5\u03c4\u03c1\u03af\u03b1\u03c3\u03b7 \u03ba\u03b1\u03b9 \u03c4\u03b7\u03bd \u03b1\u03bd\u03c4\u03b9\u03bc\u03b5\u03c4\u03ce\u03c0\u03b9\u03c3\u03b7 \u03b1\u03c5\u03c4\u03ce\u03bd \u03c4\u03c9\u03bd \u03bc\u03b5\u03bb\u03bb\u03bf\u03bd\u03c4\u03b9\u03ba\u03ce\u03bd \u03ba\u03c1\u03af\u03c3\u03b5\u03c9\u03bd.<\/p>\n

\u039f \u03bd\u03ad\u03bf\u03c2 \u03b9\u03cc\u03c2 SARS-CoV-2 \u03b1\u03bd\u03ae\u03ba\u03b5\u03b9 \u03c3\u03c4\u03b7\u03bd \u03bf\u03b9\u03ba\u03bf\u03b3\u03ad\u03bd\u03b5\u03b9\u03b1 Coronaviridae, \u03c6\u03ad\u03c1\u03b5\u03b9 \u03bc\u03bf\u03bd\u03cc\u03ba\u03bb\u03c9\u03bd\u03bf RNA (ssRNA) \u03c9\u03c2 \u03b3\u03b5\u03bd\u03b5\u03c4\u03b9\u03ba\u03cc \u03c5\u03bb\u03b9\u03ba\u03cc \u03ba\u03b1\u03b9 \u03ad\u03c7\u03b5\u03b9 \u03bc\u03b5\u03b3\u03ac\u03bb\u03b7 \u03bf\u03bc\u03bf\u03b9\u03cc\u03c4\u03b7\u03c4\u03b1 \u03bc\u03b5 \u03c4\u03bf\u03bd SARS-CoV, \u03b1\u03c6\u03bf\u03cd \u03b5\u03bc\u03c6\u03b1\u03bd\u03af\u03b6\u03b5\u03b9 \u03b3\u03b5\u03bd\u03b5\u03c4\u03b9\u03ba\u03ae \u03bf\u03bc\u03bf\u03b9\u03cc\u03c4\u03b7\u03c4\u03b1 \u03bc\u03b5 \u03b1\u03c5\u03c4\u03cc\u03bd (80% identity). \u0391\u03bb\u03bb\u03ac \u03bf \u03c0\u03b9\u03bf \u03ba\u03bf\u03bd\u03c4\u03b9\u03bd\u03cc\u03c2 \u03c3\u03c5\u03b3\u03b3\u03b5\u03bd\u03b9\u03ba\u03cc\u03c2 \u03b9\u03cc\u03c2 \u03c4\u03bf\u03c5 \u03bd\u03ad\u03bf\u03c5 \u03b9\u03bf\u03cd \u03c6\u03b1\u03af\u03bd\u03b5\u03c4\u03b1\u03b9 \u03bd\u03b1 \u03b5\u03af\u03bd\u03b1\u03b9 \u03bf \u03b9\u03cc\u03c2 \u03c4\u03c9\u03bd \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03c9\u03bd RaTG13-2013 (96% identity) \u03c3\u03cd\u03bc\u03c6\u03c9\u03bd\u03b1 \u03bc\u03b5 \u03c4\u03bf\u03c5\u03c2 Weston & Frieman (2020).<\/p>\n

\u039f\u03b9 \u03c0\u03c1\u03ce\u03c4\u03b5\u03c2 \u03bc\u03b5\u03bb\u03ad\u03c4\u03b5\u03c2 \u03c4\u03bf\u03c5 SARS-CoV-2 \u03c4\u03bf 2020 \u03c0\u03c1\u03bf\u03c3\u03b4\u03b9\u03cc\u03c1\u03b9\u03c3\u03b1\u03bd \u03cc\u03c4\u03b9 \u03bf \u03ba\u03c5\u03c4\u03c4\u03b1\u03c1\u03b9\u03ba\u03cc\u03c2 \u03c5\u03c0\u03bf\u03b4\u03bf\u03c7\u03ad\u03b1\u03c2 \u03c4\u03bf\u03c5 \u03b5\u03af\u03bd\u03b1\u03b9 \u03c4\u03bf \u03ad\u03bd\u03b6\u03c5\u03bc\u03bf ACE2, \u03cc\u03c0\u03c9\u03c2 \u03ba\u03b1\u03b9 \u03c4\u03bf\u03c5 SARS-CoV. \u039f\u03b9 \u03bc\u03b5\u03c4\u03b1\u03bb\u03bb\u03ac\u03be\u03b5\u03b9\u03c2 \u03c3\u03c4\u03bf\u03bd \u03c4\u03bf\u03bc\u03ad\u03b1 \u03b4\u03ad\u03c3\u03bc\u03b5\u03c5\u03c3\u03b7\u03c2 \u03c5\u03c0\u03bf\u03b4\u03bf\u03c7\u03ad\u03b1 (receptor-binding domain, RBD) \u03b5\u03af\u03bd\u03b1\u03b9 \u03c3\u03c5\u03c7\u03bd\u03ad\u03c2 \u03c3\u03c4\u03bf\u03c5\u03c2 \u03b9\u03bf\u03cd\u03c2 \u03b1\u03c5\u03c4\u03bf\u03cd\u03c2 \u03ba\u03b1\u03b9 \u03c6\u03b1\u03af\u03bd\u03b5\u03c4\u03b1\u03b9 \u03cc\u03c4\u03b9 \u03bf SARS-CoV-2 \u03c6\u03ad\u03c1\u03b5\u03b9 RBD \u03bc\u03b5 \u03c5\u03c8\u03b7\u03bb\u03ae \u03c3\u03c5\u03b3\u03b3\u03ad\u03bd\u03b5\u03b9\u03b1 \u03bc\u03b5 \u03c4\u03bf \u03ad\u03bd\u03b6\u03c5\u03bc\u03bf ACE2 \u03b1\u03bd\u03b8\u03c1\u03ce\u03c0\u03c9\u03bd, \u03b3\u03ac\u03c4\u03b1\u03c2, \u03ba\u03bf\u03c5\u03bd\u03b1\u03b2\u03b9\u03bf\u03cd \u03ba\u03b1\u03b9 \u03ac\u03bb\u03bb\u03c9\u03bd \u03b5\u03b9\u03b4\u03ce\u03bd \u03bc\u03b5 \u03c5\u03c8\u03b7\u03bb\u03ae \u03bf\u03bc\u03bf\u03bb\u03bf\u03b3\u03af\u03b1 \u03c5\u03c0\u03bf\u03b4\u03bf\u03c7\u03ad\u03b1 (Andersen et al, 2020). \u03a4\u03b9\u03c2 \u03c4\u03b5\u03bb\u03b5\u03c5\u03c4\u03b1\u03af\u03b5\u03c2 \u03bc\u03ad\u03c1\u03b5\u03c2 \u03c0\u03b9\u03c3\u03c4\u03bf\u03c0\u03bf\u03b9\u03ae\u03b8\u03b7\u03ba\u03b5 \u03b7 \u03bc\u03cc\u03bb\u03c5\u03bd\u03c3\u03b7 \u03b1\u03c0\u03cc \u03b1\u03c5\u03c4\u03cc\u03bd \u03c4\u03bf\u03bd \u03b9\u03cc, \u03c3\u03b5 \u03b3\u03ac\u03c4\u03b5\u03c2 \u03ba\u03b1\u03b9 \u03bc\u03b9\u03b1 \u03c4\u03af\u03b3\u03c1\u03b7, \u03b6\u03ce\u03b1 \u03c4\u03b7\u03c2 \u03af\u03b4\u03b9\u03b1\u03c2 \u03bf\u03b9\u03ba\u03bf\u03b3\u03b5\u03bd\u03b5\u03af\u03b1\u03c2 \u03c4\u03c9\u03bd \u0391\u03b9\u03bb\u03bf\u03c5\u03c1\u03b9\u03b4\u03ce\u03bd. \u03a3\u03c4\u03bf \u03c0\u03b1\u03c1\u03b5\u03bb\u03b8\u03cc\u03bd \u03c0\u03bf\u03bb\u03bb\u03bf\u03af \u03b5\u03c1\u03b5\u03c5\u03bd\u03b7\u03c4\u03ad\u03c2 \u03b5\u03af\u03c7\u03b1\u03bd \u03b5\u03c0\u03b9\u03c3\u03b7\u03bc\u03ac\u03bd\u03b5\u03b9 \u03cc\u03c4\u03b9 \u03b4\u03b5\u03bd \u03ad\u03c7\u03b5\u03b9 \u03bc\u03b5\u03bb\u03b5\u03c4\u03b7\u03b8\u03b5\u03af \u03b9\u03ba\u03b1\u03bd\u03bf\u03c0\u03bf\u03b9\u03b7\u03c4\u03b9\u03ba\u03ac \u03c4\u03bf \u03b9\u03ca\u03ba\u03cc \u03c6\u03bf\u03c1\u03c4\u03af\u03bf \u03c3\u03b5 \u03c0\u03bf\u03bb\u03bb\u03ac \u03b5\u03af\u03b4\u03b7 \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03c9\u03bd \u03b1\u03bb\u03bb\u03ac \u03ba\u03b1\u03b9 \u03b8\u03b7\u03bb\u03b1\u03c3\u03c4\u03b9\u03ba\u03ce\u03bd (\u03c7\u03bf\u03af\u03c1\u03bf\u03b9, \u03ac\u03bb\u03bf\u03b3\u03b1, \u03b5\u03c1\u03af\u03c6\u03b9\u03b1, \u03c3\u03ba\u03b1\u03c4\u03b6\u03cc\u03c7\u03bf\u03b9\u03c1\u03bf\u03b9, \u03ba\u03bf\u03c5\u03bd\u03ac\u03b2\u03b9\u03b1, \u03ba\u03b1\u03c4\u03bf\u03b9\u03ba\u03af\u03b4\u03b9\u03b1 \u03ba\u03b1\u03b9 \u03bf\u03b9\u03ba\u03cc\u03c3\u03b9\u03c4\u03b1 \u03b6\u03ce\u03b1, \u03ba.\u03b1.) \u03ba\u03b1\u03b9 \u03ac\u03bb\u03bb\u03c9\u03bd \u03b5\u03b9\u03b4\u03ce\u03bd \u03c0\u03bf\u03c5 \u03b5\u03af\u03bd\u03b1\u03b9 \u03b5\u03bd\u03b4\u03b9\u03ac\u03bc\u03b5\u03c3\u03bf\u03b9 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ad\u03c2 \u03c4\u03c9\u03bd \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03ce\u03bd \u03c0\u03b1\u03c1\u03cc\u03c4\u03b9 \u03bf \u03ba\u03af\u03bd\u03b4\u03c5\u03bd\u03bf\u03c2 \u03bc\u03b9\u03b1\u03c2 \u03bd\u03ad\u03b1\u03c2 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b1\u03c2 \u03ae\u03c4\u03b1\u03bd \u03c0\u03b9\u03b8\u03b1\u03bd\u03cc\u03c2.<\/p>\n

\"\u038c\u03c4\u03b1\u03bd<\/a>
\u038c\u03c4\u03b1\u03bd \u03c7\u03c1\u03b7\u03c3\u03b9\u03bc\u03bf\u03c0\u03bf\u03b9\u03ae\u03b8\u03b7\u03ba\u03b5 \u03c4\u03bf \u03b3\u03bf\u03bd\u03af\u03b4\u03b9\u03bf spike \u03c4\u03bf\u03c5 WIV1-CoV \u03c3\u03b5 \u03b4\u03b9\u03b1\u03b3\u03bf\u03bd\u03b9\u03b4\u03b9\u03b1\u03ba\u03ac \u03c0\u03bf\u03bd\u03c4\u03af\u03ba\u03b9\u03b1 \u03c4\u03b1 \u03bf\u03c0\u03bf\u03af\u03b1 \u03b5\u03af\u03c7\u03b1\u03bd, \u03bc\u03b5\u03c4\u03ac \u03b1\u03c0\u03cc \u03b3\u03b5\u03bd\u03b5\u03c4\u03b9\u03ba\u03ae \u03bc\u03b5\u03c4\u03b1\u03c4\u03c1\u03bf\u03c0\u03ae, \u03c4\u03b7 \u03b4\u03c5\u03bd\u03b1\u03c4\u03cc\u03c4\u03b7\u03c4\u03b1 \u03bd\u03b1 \u03b5\u03ba\u03c6\u03c1\u03ac\u03b6\u03bf\u03c5\u03bd \u03c4\u03bf\u03bd \u03c5\u03c0\u03bf\u03b4\u03bf\u03c7\u03ad\u03b1 ACE2 \u03cc\u03c0\u03c9\u03c2 \u03c4\u03b1 \u03b1\u03bd\u03b8\u03c1\u03ce\u03c0\u03b9\u03bd\u03b1 \u03ba\u03cd\u03c4\u03c4\u03b1\u03c1\u03b1, \u03c4\u03b1 \u03c0\u03b5\u03b9\u03c1\u03ac\u03bc\u03b1\u03c4\u03b1 \u03ad\u03b4\u03b5\u03b9\u03be\u03b1\u03bd \u03cc\u03c4\u03b9 \u03bf WIV1-CoV \u03ad\u03c7\u03b5\u03b9 \u03c0\u03c1\u03ac\u03b3\u03bc\u03b1\u03c4\u03b9 \u03bc\u03b5\u03b3\u03ac\u03bb\u03b7 \u03c0\u03b9\u03b8\u03b1\u03bd\u03cc\u03c4\u03b7\u03c4\u03b1 \u03b5\u03bc\u03c6\u03ac\u03bd\u03b9\u03c3\u03b7\u03c2 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03b9\u03ce\u03bd \u03c3\u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf \u03ba\u03b1\u03b9 \u03c4\u03b7\u03bd \u03bc\u03b5\u03c4\u03ac\u03b4\u03bf\u03c3\u03b7 \u03b1\u03bd\u03ac\u03bc\u03b5\u03c3\u03b1 \u03c3\u03b5 \u03b1\u03bd\u03b8\u03c1\u03ce\u03c0\u03bf\u03c5\u03c2.<\/figcaption><\/figure>\n

\u03a4\u03bf 2019 \u03bc\u03b9\u03b1 \u03bc\u03b5\u03c4\u03b1\u03c0\u03ae\u03b4\u03b7\u03c3\u03b7 \u03c4\u03bf\u03c5 \u03b9\u03bf\u03cd SADS-CoV \u03b1\u03c0\u03cc \u03c4\u03b9\u03c2 \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03b5\u03c2 \u03c3\u03c4\u03bf\u03c5\u03c2 \u03c7\u03bf\u03af\u03c1\u03bf\u03c5\u03c2 \u03c0\u03b1\u03c1\u03b1\u03c4\u03b7\u03c1\u03ae\u03b8\u03b7\u03ba\u03b5 \u03b5\u03c0\u03af\u03c3\u03b7\u03c2 (Gui et al, 2019). \u039f Corman \u03ba\u03b1\u03b9 \u03bf\u03b9 \u03c3\u03c5\u03bd\u03b5\u03c1\u03b3\u03ac\u03c4\u03b5\u03c2 \u03c4\u03bf\u03c5 \u03c4\u03bf 2014, \u03b3\u03bd\u03c9\u03c1\u03af\u03b6\u03bf\u03bd\u03c4\u03b1\u03c2 \u03cc\u03c4\u03b9 \u03bf\u03b9 \u03b9\u03bf\u03af \u03c3\u03c5\u03bd\u03b5\u03be\u03b5\u03bb\u03af\u03c3\u03c3\u03bf\u03bd\u03c4\u03b1\u03b9 \u03c3\u03c4\u03bf\u03c5\u03c2 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ad\u03c2 \u03c4\u03bf\u03c5\u03c2, \u03bc\u03b5\u03bb\u03ad\u03c4\u03b7\u03c3\u03b1\u03bd \u03c4\u03b1 \u03ba\u03cc\u03c0\u03c1\u03b1\u03bd\u03b1 \u03c4\u03c9\u03bd \u03c3\u03ba\u03b1\u03c4\u03b6\u03cc\u03c7\u03bf\u03b9\u03c1\u03c9\u03bd \u03c3\u03c4\u03b7\u03bd \u0393\u03b5\u03c1\u03bc\u03b1\u03bd\u03af\u03b1 \u03c0\u03bf\u03c5 \u03b1\u03bd\u03ae\u03ba\u03bf\u03c5\u03bd \u03c3\u03c4\u03b7\u03bd \u03c0\u03b9\u03bf \u03ba\u03bf\u03bd\u03c4\u03b9\u03bd\u03ae \u03c4\u03ac\u03be\u03b7 \u03c4\u03c9\u03bd \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03c9\u03bd, \u03c4\u03b1 Eulipotyphla, \u03ba\u03b1\u03b9 \u03b2\u03c1\u03ae\u03ba\u03b1\u03bd \u03cc\u03c4\u03b9 \u03c3\u03b5 \u03c0\u03bf\u03c3\u03bf\u03c3\u03c4\u03ac \u03ac\u03bd\u03c9 60% \u03bf\u03b9 \u03c3\u03ba\u03b1\u03c4\u03b6\u03cc\u03c7\u03bf\u03b9\u03c1\u03bf\u03b9 \u03c4\u03b7\u03c2 \u0395\u03c5\u03c1\u03ce\u03c0\u03b7\u03c2, \u03bf\u03b9 Erinaceus europaeus, \u03ae\u03c4\u03b1\u03bd \u03bc\u03bf\u03bb\u03c5\u03c3\u03bc\u03ad\u03bd\u03bf\u03b9 \u03bc\u03b5 \u03ad\u03bd\u03b1\u03bd \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03cc \u03c0\u03bf\u03bb\u03cd \u03cc\u03bc\u03bf\u03b9\u03bf \u03bc\u03b5 \u03c4\u03bf\u03bd MERS CoV.<\/p>\n

\u0397 \u03b2\u03b9\u03bf\u03bb\u03bf\u03b3\u03b9\u03ba\u03ae \u03ad\u03c1\u03b5\u03c5\u03bd\u03b1 \u03bb\u03bf\u03b9\u03c0\u03cc\u03bd, \u03b1\u03bd \u03ba\u03b1\u03b9 \u03c0\u03b5\u03c1\u03b9\u03bf\u03c1\u03b9\u03c3\u03bc\u03ad\u03bd\u03b7 \u03c0\u03c1\u03b9\u03bd \u03c4\u03bf 2019, \u03ad\u03b4\u03b5\u03b9\u03be\u03b5 \u03c4\u03cc\u03c3\u03bf \u03c4\u03b7\u03bd \u03bc\u03b5\u03b3\u03ac\u03bb\u03b7 \u03c0\u03b9\u03b8\u03b1\u03bd\u03cc\u03c4\u03b7\u03c4\u03b1 \u03bc\u03b5\u03c4\u03b1\u03bb\u03bb\u03b1\u03b3\u03ae\u03c2 \u03ba\u03b1\u03b9 \u03b4\u03b7\u03bc\u03b9\u03bf\u03c5\u03c1\u03b3\u03af\u03b1\u03c2 \u03b5\u03bd\u03cc\u03c2 \u03c0\u03b9\u03bf \u03b5\u03c0\u03b9\u03ba\u03af\u03bd\u03b4\u03c5\u03bd\u03bf\u03c5 \u03b9\u03bf\u03cd, \u03b1\u03bb\u03bb\u03ac \u03ba\u03b1\u03b9 \u03c4\u03b7\u03bd \u03b5\u03bb\u03bb\u03b9\u03c0\u03ae \u03b3\u03bd\u03ce\u03c3\u03b7 \u03c4\u03b7\u03c2 \u03b4\u03b9\u03b1\u03b5\u03b9\u03b4\u03b9\u03ba\u03ae\u03c2 \u03ba\u03b1\u03b9 \u03b5\u03bd\u03b4\u03bf\u03b5\u03b9\u03b4\u03b9\u03ba\u03ae\u03c2 \u03bc\u03b5\u03c4\u03ac\u03b4\u03bf\u03c3\u03b7\u03c2 \u03c4\u03c9\u03bd \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03ce\u03bd \u03c3\u03b5 \u03b4\u03b9\u03b1\u03c6\u03bf\u03c1\u03b5\u03c4\u03b9\u03ba\u03ac \u03b5\u03af\u03b4\u03b7, \u03b5\u03af\u03c4\u03b5 \u03b1\u03c5\u03c4\u03ac \u03b5\u03af\u03bd\u03b1\u03b9 \u03bf\u03b9\u03ba\u03cc\u03c3\u03b9\u03c4\u03b1, \u03b5\u03af\u03c4\u03b5 \u03b5\u03af\u03bd\u03b1\u03b9 \u03b5\u03b4\u03ce\u03b4\u03b9\u03bc\u03b1, \u03b5\u03af\u03c4\u03b5 \u03b6\u03bf\u03c5\u03bd \u03c3\u03c4\u03b7 \u03c6\u03cd\u03c3\u03b7. \u0395\u03ba\u03c4\u03cc\u03c2 \u03b1\u03c0\u03cc \u03c4\u03bf\u03bd MERS-CoV \u03ba\u03b1\u03bd\u03ad\u03bd\u03b1\u03c2 \u03ac\u03bb\u03bb\u03bf\u03c2 \u03b9\u03cc\u03c2 \u03b4\u03b5\u03bd \u03b9\u03c7\u03bd\u03b7\u03bb\u03b1\u03c4\u03ae\u03b8\u03b7\u03ba\u03b5 \u03bf\u03bb\u03bf\u03ba\u03bb\u03b7\u03c1\u03c9\u03bc\u03ad\u03bd\u03b1 \u03c9\u03c2 \u03c0\u03c1\u03bf\u03c2 \u03c4\u03bf\u03bd \u03ba\u03cd\u03c1\u03b9\u03bf \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ae \u03c4\u03bf\u03c5 \u03ba\u03b1\u03b9 \u03b1\u03c5\u03c4\u03cc \u03b4\u03b7\u03bc\u03b9\u03bf\u03c5\u03c1\u03b3\u03b5\u03af \u03bc\u03b5\u03b3\u03ac\u03bb\u03b5\u03c2 \u03b1\u03b2\u03b5\u03b2\u03b1\u03b9\u03cc\u03c4\u03b7\u03c4\u03b5\u03c2 \u03c9\u03c2 \u03c0\u03c1\u03bf\u03c2 \u03c4\u03bf\u03bd \u03ad\u03bb\u03b5\u03b3\u03c7\u03bf \u03c4\u03b7\u03c2 \u03bd\u03cc\u03c3\u03bf\u03c5 \u03c0\u03bf\u03c5 \u03c0\u03c1\u03bf\u03ba\u03b1\u03bb\u03b5\u03af.<\/strong><\/p>\n

\u0397 \u03b5\u03be\u03b5\u03bb\u03b9\u03c3\u03c3\u03cc\u03bc\u03b5\u03bd\u03b7 \u03bc\u03b5 \u03c5\u03c8\u03b7\u03bb\u03bf\u03cd\u03c2 \u03c1\u03c5\u03b8\u03bc\u03bf\u03cd\u03c2 \u03c0\u03b1\u03bd\u03b4\u03b7\u03bc\u03af\u03b1 \u03b3\u03b5\u03bd\u03bd\u03ac \u03bd\u03ad\u03b1 \u03b5\u03c0\u03b9\u03c3\u03c4\u03b7\u03bc\u03bf\u03bd\u03b9\u03ba\u03ac \u03b5\u03c1\u03c9\u03c4\u03ae\u03bc\u03b1\u03c4\u03b1 \u03ba\u03b1\u03b9 \u03b4\u03b9\u03b1\u03bc\u03bf\u03c1\u03c6\u03ce\u03bd\u03b5\u03b9 \u03bd\u03ad\u03b1 \u03c0\u03b5\u03b4\u03af\u03b1 \u03ad\u03c1\u03b5\u03c5\u03bd\u03b1\u03c2. \u0395\u03c0\u03ad\u03c6\u03b5\u03c1\u03b5 \u03c3\u03b5 \u03c0\u03b1\u03b3\u03ba\u03cc\u03c3\u03bc\u03b9\u03bf \u03b5\u03c0\u03af\u03c0\u03b5\u03b4\u03bf, \u03bc\u03ad\u03c4\u03c1\u03b1 \u03c0\u03b5\u03c1\u03b9\u03bf\u03c1\u03b9\u03c3\u03bc\u03bf\u03cd \u03c4\u03b7\u03c2 \u03b5\u03bb\u03b5\u03c5\u03b8\u03b5\u03c1\u03af\u03b1\u03c2 \u03c4\u03c9\u03bd \u03c0\u03bf\u03bb\u03b9\u03c4\u03ce\u03bd \u03c0\u03c1\u03c9\u03c4\u03bf\u03c6\u03b1\u03bd\u03ae \u03b9\u03b4\u03af\u03c9\u03c2 \u03b3\u03b9\u03b1 \u03c4\u03b7 \u0394\u03cd\u03c3\u03b7, \u03bc\u03b5 \u03bc\u03b9\u03ba\u03c1\u03ae \u03c0\u03c1\u03bf\u03c2 \u03c4\u03bf \u03c0\u03b1\u03c1\u03cc\u03bd \u03b1\u03c0\u03bf\u03c4\u03b5\u03bb\u03b5\u03c3\u03bc\u03b1\u03c4\u03b9\u03ba\u03cc\u03c4\u03b7\u03c4\u03b1. \u03a4\u03b1 \u03bc\u03ad\u03c4\u03c1\u03b1 \u03c0\u03b5\u03c1\u03b9\u03bf\u03c1\u03b9\u03c3\u03bc\u03bf\u03cd \u03ba\u03b1\u03b9 \u03ba\u03bf\u03b9\u03bd\u03c9\u03bd\u03b9\u03ba\u03ae\u03c2 \u03b1\u03c0\u03bf\u03bc\u03cc\u03bd\u03c9\u03c3\u03b7\u03c2 \u03b5\u03af\u03bd\u03b1\u03b9 \u03bc\u03ad\u03c4\u03c1\u03b1 \u03b1\u03bd\u03c4\u03b9\u03bc\u03b5\u03c4\u03ce\u03c0\u03b9\u03c3\u03b7\u03c2 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03b9\u03ce\u03bd \u03ba\u03b1\u03b9 \u03c3\u03b5 \u03c0\u03b5\u03c1\u03b1\u03c3\u03bc\u03ad\u03bd\u03bf\u03c5\u03c2 \u03b1\u03b9\u03ce\u03bd\u03b5\u03c2 \u03c3\u03c4\u03b7 \u03c7\u03ce\u03c1\u03b1 (*1).<\/p>\n

\u03a3\u03c4\u03b7\u03bd \u0395\u03bb\u03bb\u03ac\u03b4\u03b1 \u03b4\u03b5\u03bd \u03ad\u03b3\u03b9\u03bd\u03b1\u03bd \u03b9\u03bf\u03bb\u03bf\u03b3\u03b9\u03ba\u03ad\u03c2 \u03bc\u03b5\u03bb\u03ad\u03c4\u03b5\u03c2 \u03b3\u03b9\u03b1 \u03c4\u03bf\u03c5\u03c2 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03cd\u03c2 \u03bf\u03cd\u03c4\u03b5 \u03bc\u03b5\u03bb\u03ad\u03c4\u03b5\u03c2 \u03b5\u03bd\u03c4\u03bf\u03c0\u03b9\u03c3\u03bc\u03bf\u03cd \u03c4\u03bf\u03c5\u03c2 \u03c3\u03c4\u03b7\u03bd \u03c0\u03b1\u03bd\u03af\u03b4\u03b1, \u03ad\u03c4\u03c3\u03b9 \u03ce\u03c3\u03c4\u03b5 \u03bd\u03b1 \u03b9\u03c7\u03bd\u03b7\u03bb\u03b1\u03c4\u03b7\u03b8\u03b5\u03af \u03bf \u03b1\u03bd\u03b1\u03bc\u03b5\u03bd\u03cc\u03bc\u03b5\u03bd\u03bf\u03c2 \u03ba\u03af\u03bd\u03b4\u03c5\u03bd\u03bf\u03c2 \u03bc\u03b9\u03b1\u03c2 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b1\u03c2, \u03c0\u03ad\u03c1\u03b1\u03bd \u03c4\u03bf\u03c5 \u03b5\u03bd\u03c4\u03bf\u03c0\u03b9\u03c3\u03bc\u03bf\u03cd \u03b1\u03bd\u03b8\u03c1\u03ce\u03c0\u03c9\u03bd (\u03c4\u03c9\u03bd \u03ba\u03c1\u03bf\u03c5\u03c3\u03bc\u03ac\u03c4\u03c9\u03bd) \u03c0\u03bf\u03c5 \u03b3\u03af\u03bd\u03b5\u03c4\u03b1\u03b9 \u03bc\u03b5\u03c3\u03bf\u03cd\u03c3\u03b7\u03c2 \u03c4\u03b7\u03c2 \u03c0\u03b1\u03bd\u03b4\u03b7\u03bc\u03af\u03b1\u03c2, \u03ad\u03c4\u03c3\u03b9 \u03ce\u03c3\u03c4\u03b5 \u03bd\u03b1 \u03c5\u03c0\u03ac\u03c1\u03c7\u03bf\u03c5\u03bd \u03b4\u03b5\u03b4\u03bf\u03bc\u03ad\u03bd\u03b1 \u03b3\u03b9\u03b1 \u03c4\u03bf\u03bd \u03bf\u03bb\u03bf\u03ba\u03bb\u03b7\u03c1\u03c9\u03bc\u03ad\u03bd\u03bf \u03ad\u03bb\u03b5\u03b3\u03c7\u03bf \u03c4\u03b7\u03c2 \u03b4\u03b9\u03b1\u03c3\u03c0\u03bf\u03c1\u03ac\u03c2 \u03c4\u03bf\u03c5 \u03ba\u03b1\u03b9 \u03b4\u03b5\u03b4\u03bf\u03bc\u03ad\u03bd\u03b1 \u03b3\u03b9\u03b1 \u03c4\u03b7\u03bd \u03c0\u03b5\u03c1\u03b9\u03bf\u03b4\u03b9\u03ba\u03ae \u03ae \u03bc\u03b7 \u03b5\u03c0\u03b1\u03bd\u03b5\u03bc\u03c6\u03ac\u03bd\u03b9\u03c3\u03b7 \u03c4\u03bf\u03c5. \u039c\u03b9\u03b1 \u03bf\u03b9\u03ba\u03bf\u03bb\u03bf\u03b3\u03b9\u03ba\u03ae \u03c0\u03c1\u03bf\u03c3\u03ad\u03b3\u03b3\u03b9\u03c3\u03b7 \u03b8\u03b1 \u03bc\u03c0\u03bf\u03c1\u03bf\u03cd\u03c3\u03b5 \u03bd\u03b1 \u03b4\u03ce\u03c3\u03b5\u03b9 \u03c0\u03bb\u03b7\u03c1\u03bf\u03c6\u03bf\u03c1\u03af\u03b5\u03c2 \u03c7\u03c1\u03ae\u03c3\u03b9\u03bc\u03b5\u03c2 \u03b3\u03b9\u03b1 \u03bc\u03b9\u03b1 \u03c4\u03ad\u03c4\u03bf\u03b9\u03b1 \u03c0\u03c1\u03cc\u03b2\u03bb\u03b5\u03c8\u03b7.<\/p>\n

\u03a4\u03bf \u03ba\u03b5\u03bd\u03c4\u03c1\u03b9\u03ba\u03cc \u03b5\u03c1\u03ce\u03c4\u03b7\u03bc\u03b1 \u03bb\u03bf\u03b9\u03c0\u03cc\u03bd, \u03c4\u03bf\u03c5 \u03c0\u03c1\u03bf\u03c3\u03b4\u03b9\u03bf\u03c1\u03b9\u03c3\u03bc\u03bf\u03cd \u03c4\u03c9\u03bd \u03b5\u03bd\u03b4\u03b9\u03ac\u03bc\u03b5\u03c3\u03c9\u03bd \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ce\u03bd \u03c3\u03b5 \u03cc\u03bb\u03b5\u03c2 \u03c4\u03b9\u03c2 \u03c6\u03bf\u03bd\u03b9\u03ba\u03ad\u03c2 \u03c0\u03b1\u03bd\u03b4\u03b7\u03bc\u03af\u03b5\u03c2 \u03b6\u03c9\u03bf\u03bd\u03bf\u03c4\u03b9\u03ba\u03ce\u03bd \u03c0\u03b1\u03b8\u03bf\u03b3\u03cc\u03bd\u03c9\u03bd, \u03b5\u03af\u03bd\u03b1\u03b9 \u03c4\u03bf \u03ba\u03c1\u03af\u03c3\u03b9\u03bc\u03bf \u03b5\u03c1\u03ce\u03c4\u03b7\u03bc\u03b1 \u03ba\u03b1\u03b9 \u03c3\u03b5 \u03b1\u03c5\u03c4\u03ae\u03bd \u03c4\u03b7\u03bd \u03c0\u03b1\u03bd\u03b4\u03b7\u03bc\u03af\u03b1. \u039f\u03b9 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ad\u03c2 \u03b5\u03af\u03bd\u03b1\u03b9 \u03bf\u03b9 \u03b1\u03c0\u03bf\u03b8\u03ae\u03ba\u03b5\u03c2 \u03c4\u03bf\u03c5 \u03b9\u03bf\u03cd \u03c0\u03bf\u03c5 \u03c0\u03b1\u03c1\u03b1\u03bc\u03ad\u03bd\u03bf\u03c5\u03bd \u03c3\u03c4\u03b7 \u03c6\u03cd\u03c3\u03b7, \u03b4\u03b9\u03b1\u03c3\u03c0\u03b5\u03af\u03c1\u03bf\u03bd\u03c4\u03b1\u03b9 \u03ba\u03b1\u03b9 \u03bc\u03b5\u03c4\u03b1\u03bb\u03bb\u03ac\u03c3\u03c3\u03bf\u03bd\u03c4\u03b1\u03b9. \u0391\u03c5\u03c4\u03cc \u03c4\u03bf \u03b3\u03b5\u03b3\u03bf\u03bd\u03cc\u03c2 \u03b5\u03af\u03bd\u03b1\u03b9 \u03ba\u03b1\u03b9 \u03bf \u03bb\u03cc\u03b3\u03bf\u03c2 \u03c0\u03bf\u03c5 \u03bf\u03b9 \u03b6\u03c9\u03bf\u03bd\u03cc\u03c3\u03bf\u03b9 \u03b4\u03b5\u03bd \u03b5\u03be\u03b1\u03bb\u03b5\u03af\u03c6\u03bf\u03bd\u03c4\u03b1\u03b9, \u03cc\u03c3\u03b1 \u03ba\u03b9 \u03b1\u03bd \u03b3\u03bd\u03c9\u03c1\u03af\u03b6\u03bf\u03c5\u03bc\u03b5 \u03b3\u03b9\u03b1 \u03c4\u03b7\u03bd \u03bc\u03bf\u03c1\u03c6\u03ae \u03ba\u03b1\u03b9 \u03c4\u03b7\u03bd \u03b2\u03b9\u03bf\u03bb\u03bf\u03b3\u03af\u03b1 \u03c4\u03bf\u03c5\u03c2 \u03ae \u03b1\u03ba\u03cc\u03bc\u03b1 \u03b3\u03b9\u03b1 \u03c4\u03b1 \u03c6\u03ac\u03c1\u03bc\u03b1\u03ba\u03b1 \u03ae \u03c4\u03b1 \u03b5\u03bc\u03b2\u03cc\u03bb\u03b9\u03b1. \u039f\u03b9 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b5\u03c2 \u03bc\u03b5\u03c4\u03c1\u03b9\u03ac\u03b6\u03bf\u03bd\u03c4\u03b1\u03b9, \u03b5\u03bb\u03ad\u03b3\u03c7\u03bf\u03bd\u03c4\u03b1\u03b9 \u03b1\u03bb\u03bb\u03ac \u03b4\u03b5\u03bd \u03b5\u03be\u03b1\u03bb\u03b5\u03af\u03c6\u03bf\u03bd\u03c4\u03b1\u03b9 \u03b5\u03c6\u03cc\u03c3\u03bf\u03bd \u03bf\u03b9 \u03b1\u03c0\u03bf\u03b8\u03ae\u03ba\u03b5\u03c2 \u03c4\u03c9\u03bd \u03c0\u03b1\u03b8\u03bf\u03b3\u03cc\u03bd\u03c9\u03bd \u03b9\u03ce\u03bd \u03ae \u03bc\u03b9\u03ba\u03c1\u03bf\u03b2\u03af\u03c9\u03bd \u03c3\u03c4\u03b7 \u03c6\u03cd\u03c3\u03b7 \u03b4\u03b9\u03b1\u03b9\u03c9\u03bd\u03af\u03b6\u03bf\u03bd\u03c4\u03b1\u03b9 \u03bc\u03ad\u03c3\u03c9 \u03c4\u03c9\u03bd \u03b6\u03ce\u03c9\u03bd \u03cc\u03c0\u03bf\u03c5 \u03ba\u03b1\u03b9 \u03bc\u03b5\u03c4\u03b1\u03bb\u03bb\u03ac\u03c3\u03c3\u03bf\u03bd\u03c4\u03b1\u03b9 \u03b4\u03b7\u03bc\u03b9\u03bf\u03c5\u03c1\u03b3\u03ce\u03bd\u03c4\u03b1\u03c2 \u03bd\u03ad\u03b1 \u03c0\u03b9\u03bf \u03bc\u03bf\u03bb\u03c5\u03c3\u03bc\u03b1\u03c4\u03b9\u03ba\u03ac \u03ae \u03b8\u03b1\u03bd\u03b1\u03c4\u03b7\u03c6\u03cc\u03c1\u03b1 \u03c3\u03c4\u03b5\u03bb\u03ad\u03c7\u03b7. \u039f MERS-CoV \u03b1\u03bd\u03c4\u03b9\u03bc\u03b5\u03c4\u03c9\u03c0\u03af\u03c3\u03c4\u03b7\u03ba\u03b5 \u03b1\u03ba\u03c1\u03b9\u03b2\u03ce\u03c2, \u03b5\u03c0\u03b5\u03b9\u03b4\u03ae \u03c0\u03c1\u03bf\u03c3\u03b4\u03b9\u03bf\u03c1\u03af\u03c3\u03c4\u03b7\u03ba\u03b5 \u03b5\u03cd\u03ba\u03bf\u03bb\u03b1 \u03bf \u03b5\u03bd\u03b4\u03b9\u03ac\u03bc\u03b5\u03c3\u03bf\u03c2 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ae\u03c2 \u03c4\u03bf\u03c5. \u0388\u03bd\u03b1 \u03b5\u03bc\u03b2\u03cc\u03bb\u03b9\u03bf \u03c3\u03b5 \u03b6\u03ce\u03b1 \u03b5\u03af\u03bd\u03b1\u03b9 \u03c0\u03ac\u03bd\u03c4\u03b1 \u03c0\u03b9\u03bf \u03b5\u03cd\u03ba\u03bf\u03bb\u03bf \u03ba\u03b1\u03b9 \u03b1\u03ba\u03af\u03bd\u03b4\u03c5\u03bd\u03bf \u03bd\u03b1 \u03b5\u03c6\u03b1\u03c1\u03bc\u03bf\u03c3\u03c4\u03b5\u03af, \u03b5\u03bd\u03ce \u03c0\u03b5\u03c1\u03b9\u03bf\u03c1\u03af\u03b6\u03b5\u03b9 \u03b4\u03c1\u03b1\u03c3\u03c4\u03b9\u03ba\u03ac \u03c4\u03b7\u03bd \u03b5\u03be\u03ac\u03c0\u03bb\u03c9\u03c3\u03b7 \u03c4\u03bf\u03c5 \u03b9\u03bf\u03cd \u03ae \u03c4\u03b7\u03bd \u03bc\u03b7\u03b4\u03b5\u03bd\u03af\u03b6\u03b5\u03b9 \u03c0\u03c1\u03b1\u03ba\u03c4\u03b9\u03ba\u03ac. T\u03b1 \u03bc\u03ad\u03c4\u03c1\u03b1 \u03b1\u03c0\u03bf\u03bc\u03cc\u03bd\u03c9\u03c3\u03b7\u03c2 \u03ba\u03b1\u03b9 \u03c0\u03b5\u03c1\u03b9\u03bf\u03c1\u03b9\u03c3\u03bc\u03bf\u03cd \u03c4\u03c9\u03bd \u03bb\u03b1\u03ce\u03bd \u03c0\u03bf\u03c5 \u03b5\u03c6\u03b1\u03c1\u03bc\u03cc\u03b6\u03bf\u03bd\u03c4\u03b1\u03b9 \u03b3\u03b9\u03b1 \u03b1\u03b9\u03ce\u03bd\u03b5\u03c2 \u03c3\u03c4\u03b9\u03c2 \u03c0\u03b1\u03bd\u03b4\u03b7\u03bc\u03af\u03b5\u03c2, \u03bc\u03b5\u03c4\u03c1\u03b9\u03ac\u03b6\u03bf\u03c5\u03bd \u03c4\u03bf \u03c0\u03c1\u03cc\u03b2\u03bb\u03b7\u03bc\u03b1 \u03c7\u03c9\u03c1\u03af\u03c2 \u03bd\u03b1 \u03b5\u03be\u03b1\u03bb\u03b5\u03af\u03c6\u03bf\u03c5\u03bd \u03c4\u03bf \u03b1\u03af\u03c4\u03b9\u03bf.<\/strong><\/p>\n

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\u03b5\u03af\u03bd\u03b1\u03b9 \u03b5\u03c0\u03b9\u03b2\u03b5\u03b2\u03bb\u03b7\u03bc\u03ad\u03bd\u03b7 \u03ba\u03b1\u03b9 \u03b3\u03b9\u03b1 \u03c4\u03b7\u03bd \u03ad\u03b3\u03ba\u03b1\u03b9\u03c1\u03b7 \u03c0\u03c1\u03cc\u03b3\u03bd\u03c9\u03c3\u03b7 \u03bd\u03ad\u03c9\u03bd \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03b9\u03ce\u03bd \u03ba\u03b1\u03b9 \u03b3\u03b9\u03b1 \u03c4\u03bf\u03bd \u03ad\u03bb\u03b5\u03b3\u03c7\u03cc \u03c4\u03bf\u03c5\u03c2. \u039f\u03b9 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03af \u03b4\u03b5\u03bd \u03b1\u03bd\u03c4\u03b9\u03bc\u03b5\u03c4\u03c9\u03c0\u03af\u03b6\u03bf\u03bd\u03c4\u03b1\u03b9 \u03c7\u03c9\u03c1\u03af\u03c2 \u03c4\u03b7\u03bd \u03bc\u03b5\u03bb\u03ad\u03c4\u03b7 \u03c4\u03b7\u03c2 \u03b2\u03b9\u03bf\u03b3\u03b5\u03c9\u03b3\u03c1\u03b1\u03c6\u03af\u03b1\u03c2 \u03ba\u03b1\u03b9 \u03c4\u03bf\u03c5 \u03c0\u03bb\u03ad\u03b3\u03bc\u03b1\u03c4\u03bf\u03c2 \u03c0\u03b1\u03b8\u03bf\u03b3\u03cc\u03bd\u03bf\u03c5 \u03ba\u03b1\u03b9 \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ae\/\u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ce\u03bd. \u039f\u03b9 \u03b3\u03bd\u03c9\u03c3\u03c4\u03bf\u03af \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03af \u03c0\u03c1\u03bf\u03c3\u03b2\u03ac\u03bb\u03bb\u03bf\u03c5\u03bd \u03c0\u03bf\u03bb\u03bb\u03ac \u03cc\u03c1\u03b3\u03b1\u03bd\u03b1 \u03ba\u03b1\u03b9 \u03bc\u03c0\u03bf\u03c1\u03b5\u03af \u03bd\u03b1 \u03c0\u03c1\u03bf\u03ba\u03b1\u03bb\u03ad\u03c3\u03bf\u03c5\u03bd \u03c0\u03bf\u03bb\u03c5\u03bf\u03c1\u03b3\u03b1\u03bd\u03b9\u03ba\u03ae \u03b1\u03bd\u03b5\u03c0\u03ac\u03c1\u03ba\u03b5\u03b9\u03b1, \u03c4\u03cc\u03c3\u03bf \u03c3\u03b5 \u03b6\u03ce\u03b1 \u03cc\u03c3\u03bf \u03ba\u03b1\u03b9 \u03c3\u03c4\u03bf\u03bd \u03ac\u03bd\u03b8\u03c1\u03c9\u03c0\u03bf. \u0395\u03c0\u03b5\u03b9\u03b4\u03ae \u03c0\u03c1\u03bf\u03c3\u03b2\u03ac\u03bb\u03bb\u03bf\u03c5\u03bd \u03c4\u03bf \u03c0\u03b5\u03c0\u03c4\u03b9\u03ba\u03cc \u03c3\u03cd\u03c3\u03c4\u03b7\u03bc\u03b1, 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\u03b5\u03c0\u03b9\u03c6\u03b1\u03bd\u03b5\u03b9\u03b1\u03ba\u03ac \u03bd\u03b5\u03c1\u03ac, \u03c4\u03bf \u03ad\u03b4\u03b1\u03c6\u03bf\u03c2 \u03b1\u03bb\u03bb\u03ac \u03ba\u03b1\u03b9 \u03c4\u03bf\u03bd \u03b1\u03ad\u03c1\u03b1. \u03a3\u03c4\u03b9\u03c2 \u03c0\u03b5\u03c1\u03b9\u03bf\u03c7\u03ad\u03c2 \u03c0\u03bf\u03c5 \u03b7 \u03c0\u03b1\u03bd\u03b4\u03b7\u03bc\u03af\u03b1 \u03ad\u03c7\u03b5\u03b9 \u03b4\u03c1\u03b1\u03bc\u03b1\u03c4\u03b9\u03ba\u03ad\u03c2 \u03b4\u03b9\u03b1\u03c3\u03c4\u03ac\u03c3\u03b5\u03b9\u03c2, \u03b5\u03af\u03bd\u03b1\u03b9 \u03c0\u03b9\u03b8\u03b1\u03bd\u03ae \u03b7 \u03bc\u03cc\u03bb\u03c5\u03bd\u03c3\u03b7 \u03c4\u03bf\u03c5 \u03c0\u03b5\u03c1\u03b9\u03b2\u03ac\u03bb\u03bb\u03bf\u03bd\u03c4\u03bf\u03c2 \u03b1\u03c0\u03cc \u03c4\u03b1 \u03bb\u03cd\u03bc\u03b1\u03c4\u03b1.<\/strong> \u0399\u03b4\u03af\u03c9\u03c2 \u03cc\u03c0\u03bf\u03c5 \u03c4\u03b1 \u03bb\u03cd\u03bc\u03b1\u03c4\u03b1 \u03c7\u03cd\u03bd\u03bf\u03bd\u03c4\u03b1\u03b9 \u03c3\u03b5 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\u03b1) \u03bc\u03ad\u03c3\u03c9 \u03c4\u03c9\u03bd \u03b2\u03b9\u03bf\u03b1\u03b5\u03c1\u03bf\u03bb\u03c5\u03bc\u03ac\u03c4\u03c9\u03bd \u03c3\u03c4\u03b9\u03c2 \u03c0\u03cc\u03bb\u03b5\u03b9\u03c2 \u03ba\u03b1\u03b9 \u03ba\u03bf\u03bd\u03c4\u03ac \u03c3\u03b5 \u03b2\u03b9\u03bf\u03bb\u03bf\u03b3\u03b9\u03ba\u03bf\u03cd\u03c2 \u03ba\u03b1\u03b8\u03b1\u03c1\u03b9\u03c3\u03bc\u03bf\u03cd\u03c2,<\/p>\n

\u03b2) \u03c3\u03c4\u03b1 \u03b8\u03b1\u03bb\u03b1\u03c3\u03c3\u03b9\u03bd\u03ac \u03ba\u03b1\u03b9 \u03b5\u03c0\u03b9\u03c6\u03b1\u03bd\u03b5\u03b9\u03b1\u03ba\u03ac \u03bd\u03b5\u03c1\u03ac \u03bb\u03cc\u03b3\u03c9 \u03c4\u03b7\u03c2 \u03bc\u03b5\u03b3\u03ac\u03bb\u03b7\u03c2 \u03c3\u03c7\u03b5\u03c4\u03b9\u03ba\u03ac \u03b1\u03bd\u03c4\u03bf\u03c7\u03ae\u03c2 \u03c4\u03bf\u03c5 \u03b9\u03bf\u03cd \u03c3\u03c4\u03bf \u03c0\u03b5\u03c1\u03b9\u03b2\u03ac\u03bb\u03bb\u03bf\u03bd.<\/p>\n

\"\u03a3\u03c4\u03bf<\/a>
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\u0395\u03c0\u03af\u03c3\u03b7\u03c2 \u03c0\u03c1\u03bf\u03b3\u03c1\u03ac\u03bc\u03bc\u03b1\u03c4\u03b1 \u03b2\u03b1\u03c3\u03b9\u03ba\u03ae\u03c2 \u03ad\u03c1\u03b5\u03c5\u03bd\u03b1\u03c2 \u03cc\u03c0\u03c9\u03c2 \u03c0.\u03c7. \u03b7 \u03c0\u03b1\u03c1\u03b1\u03ba\u03bf\u03bb\u03bf\u03cd\u03b8\u03b7\u03c3\u03b7 \u03c4\u03b7\u03c2 \u03b2\u03b9\u03bf\u03c0\u03bf\u03b9\u03ba\u03b9\u03bb\u03cc\u03c4\u03b7\u03c4\u03b1\u03c2 \u03ba\u03b1\u03b9 \u03c4\u03b7\u03c2 \u03b4\u03c5\u03bd\u03b1\u03bc\u03b9\u03ba\u03ae\u03c2 \u03c0\u03bb\u03b7\u03b8\u03c5\u03c3\u03bc\u03ce\u03bd \u03cc\u03bb\u03c9\u03bd \u03c4\u03c9\u03bd \u03b4\u03c5\u03bd\u03b7\u03c4\u03b9\u03ba\u03ac \u03be\u03b5\u03bd\u03b9\u03c3\u03c4\u03ce\u03bd \u03c4\u03c9\u03bd \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03ce\u03bd \u03c3\u03b5 \u03c0\u03b5\u03c1\u03b9\u03bf\u03c7\u03ad\u03c2 \u03ba\u03c1\u03af\u03c3\u03b9\u03bc\u03b5\u03c2 \u03cc\u03c0\u03c9\u03c2 \u03bf\u03b9 \u03c0\u03b5\u03c1\u03b9\u03b1\u03c3\u03c4\u03b9\u03ba\u03ad\u03c2. \u0395\u03c0\u03af\u03c3\u03b7\u03c2 \u03c0\u03c1\u03bf\u03b3\u03c1\u03ac\u03bc\u03bc\u03b1\u03c4\u03b1 \u03c3\u03b5 \u03b2\u03b9\u03cc\u03c4\u03bf\u03c0\u03bf\u03c5\u03c2 \u03bc\u03b9\u03ba\u03c1\u03ce\u03bd \u03b8\u03b7\u03bb\u03b1\u03c3\u03c4\u03b9\u03ba\u03ce\u03bd \u03ba\u03b1\u03b9 \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03c9\u03bd \u03ba.\u03b1. \u03b5\u03af\u03bd\u03b1\u03b9 \u03b1\u03c0\u03b1\u03c1\u03b1\u03af\u03c4\u03b7\u03c4\u03bf \u03bd\u03b1 \u03b3\u03af\u03bd\u03bf\u03c5\u03bd \u03ad\u03c1\u03b5\u03c5\u03bd\u03b5\u03c2 \u03b3\u03b9\u03b1 \u03bd\u03b1 \u03b5\u03bb\u03b5\u03b3\u03c7\u03b8\u03b5\u03af \u03ba\u03b1\u03b9 \u03bd\u03b1 \u03b1\u03bd\u03c4\u03b9\u03bc\u03b5\u03c4\u03c9\u03c0\u03b9\u03c3\u03c4\u03b5\u03af \u03b7 \u03c3\u03b7\u03bc\u03b5\u03c1\u03b9\u03bd\u03ae \u03b1\u03bb\u03bb\u03ac \u03ba\u03b1\u03b9 \u03c0\u03c1\u03bf\u03bb\u03b7\u03c6\u03b8\u03bf\u03cd\u03bd \u03bf\u03b9 \u03b5\u03c0\u03cc\u03bc\u03b5\u03bd\u03b5\u03c2 \u03ba\u03c1\u03af\u03c3\u03b5\u03b9\u03c2. \u0397 \u03b3\u03bd\u03ce\u03c3\u03b7 \u03c4\u03b7\u03c2 \u03c0\u03b1\u03bd\u03af\u03b4\u03b1\u03c2 \u03ba\u03b1\u03b9 \u03c4\u03b7\u03c2 \u03c7\u03bb\u03c9\u03c1\u03af\u03b4\u03b1\u03c2 \u03c4\u03b7\u03c2 \u03c7\u03ce\u03c1\u03b1\u03c2 \u03b5\u03af\u03bd\u03b1\u03b9 \u03b5\u03bb\u03bb\u03b9\u03c0\u03ae\u03c2 \u03ba\u03b1\u03b9 \u03b1\u03c0\u03bf\u03c3\u03c0\u03b1\u03c3\u03bc\u03b1\u03c4\u03b9\u03ba\u03ae, \u03c0\u03b1\u03c1\u03cc\u03c4\u03b9 \u03b5\u03af\u03bd\u03b1\u03b9 \u03bc\u03b9\u03b1 \u03b3\u03bd\u03ce\u03c3\u03b7 \u03c0\u03bf\u03bb\u03cd\u03c4\u03b9\u03bc\u03b7 \u03b3\u03b9\u03b1 \u03c4\u03b7\u03bd \u03b4\u03b7\u03bc\u03cc\u03c3\u03b9\u03b1 \u03c5\u03b3\u03b5\u03af\u03b1, \u03c4\u03b7\u03bd \u03bf\u03b9\u03ba\u03bf\u03bd\u03bf\u03bc\u03af\u03b1 \u03ba\u03b1\u03b9 \u03c4\u03b7\u03bd \u03b5\u03c0\u03b9\u03c3\u03c4\u03ae\u03bc\u03b7.<\/strong> \u03a3\u03b5 \u03b1\u03c5\u03c4\u03ae\u03bd \u03c4\u03b7\u03bd \u03c0\u03c1\u03bf\u03c3\u03c0\u03ac\u03b8\u03b5\u03b9\u03b1 \u03b7 \u03b5\u03c0\u03b9\u03c3\u03c4\u03ae\u03bc\u03b7 \u03c4\u03c9\u03bd \u03c0\u03bf\u03bb\u03b9\u03c4\u03ce\u03bd \u03ba\u03b1\u03b9 \u03b7 \u03c3\u03cd\u03b3\u03c7\u03c1\u03bf\u03bd\u03b7 \u03c4\u03b5\u03c7\u03bd\u03bf\u03bb\u03bf\u03b3\u03af\u03b1 \u03bc\u03c0\u03bf\u03c1\u03bf\u03cd\u03bd \u03bd\u03b1 \u03b2\u03bf\u03b7\u03b8\u03ae\u03c3\u03bf\u03c5\u03bd \u03c0\u03bf\u03bb\u03cd \u03ba\u03b1\u03b9 \u03b7 \u03c0\u03b1\u03bd\u03b4\u03b7\u03bc\u03af\u03b1 \u03b1\u03c5\u03c4\u03ae \u03bc\u03c0\u03bf\u03c1\u03b5\u03af \u03bd\u03b1 \u03b5\u03c0\u03b9\u03c4\u03b1\u03c7\u03cd\u03bd\u03b5\u03b9 \u03c4\u03ad\u03c4\u03bf\u03b9\u03b5\u03c2 \u03c0\u03c1\u03bf\u03c3\u03b5\u03b3\u03b3\u03af\u03c3\u03b5\u03b9\u03c2.\u00a0<\/em><\/p>\n

(*1) \u038c\u03c0\u03c9\u03c2 \u03c4\u03b7\u03c2 \u03c0\u03b1\u03bd\u03ce\u03bb\u03b7\u03c2 \u03c4\u03bf 1828, \u03cc\u03c4\u03b1\u03bd \u03bf \u039a\u03b1\u03c0\u03bf\u03b4\u03af\u03c3\u03c4\u03c1\u03b9\u03b1\u03c2 \u03ad\u03ba\u03bb\u03b5\u03b9\u03c3\u03b5 \u03c4\u03b9\u03c2 \u03b5\u03ba\u03ba\u03bb\u03b7\u03c3\u03af\u03b5\u03c2 \u03ba\u03b1\u03b9 \u03b5\u03c0\u03ad\u03b2\u03b1\u03bb\u03b5 \u03ba\u03b1\u03c4\u0384\u03bf\u03af\u03ba\u03bf\u03bd \u03c0\u03b5\u03c1\u03b9\u03bf\u03c1\u03b9\u03c3\u03bc\u03cc \u03ba\u03b1\u03b9 \u03b1\u03c5\u03c3\u03c4\u03b7\u03c1\u03cc \u03b1\u03c0\u03bf\u03ba\u03bb\u03b5\u03b9\u03c3\u03bc\u03cc \u03c4\u03b7\u03c2 \u038e\u03b4\u03c1\u03b1\u03c2 \u03ba\u03b1\u03b9 \u03c4\u03c9\u03bd \u03a3\u03c0\u03b5\u03c4\u03c3\u03ce\u03bd. <\/em><\/p>\n

\u039f \u0395. \u039b\u03c5\u03ba\u03bf\u03cd\u03b4\u03b7\u03c2 \u03c0\u03b5\u03c1\u03b9\u03b3\u03c1\u03ac\u03c6\u03b5\u03b9 \u03c0\u03b1\u03c1\u03cc\u03bc\u03bf\u03b9\u03b1 \u03bc\u03ad\u03c4\u03c1\u03b1 \u03b3\u03b9\u03b1 \u03c4\u03b7 \u03c7\u03bf\u03bb\u03ad\u03c1\u03b1 \u03b5\u03c0\u03af \u038c\u03b8\u03c9\u03bd\u03b1. \u0397 \u03bd\u03cc\u03c3\u03bf\u03c2 \u03ae\u03c1\u03b8\u03b5 \u03c4\u03bf 1854 \u03bc\u03b5 \u03c4\u03bf\u03bd \u03b3\u03b1\u03bb\u03bb\u03b9\u03ba\u03cc \u03c3\u03c4\u03c1\u03b1\u03c4\u03cc \u03c4\u03b7\u03c2 \u039a\u03c1\u03b9\u03bc\u03b1\u03af\u03b1\u03c2 \u03c0\u03bf\u03c5 \u03b5\u03af\u03c7\u03b5 \u03ba\u03b1\u03c4\u03b1\u03c6\u03b8\u03ac\u03c3\u03b5\u03b9 \u03c3\u03c4\u03bf\u03bd \u03a0\u03b5\u03b9\u03c1\u03b1\u03b9\u03ac \u03bc\u03ad\u03c3\u03c9 \u03bf\u03c0\u03bb\u03b9\u03c4\u03b1\u03b3\u03c9\u03b3\u03ce\u03bd \u03c0\u03bb\u03bf\u03af\u03c9\u03bd \u03c9\u03c2 \u03c3\u03c4\u03c1\u03b1\u03c4\u03cc\u03c2 \u039a\u03b1\u03c4\u03bf\u03c7\u03ae\u03c2 \u03ba\u03b1\u03b9 \u03b4\u03b9\u03b1\u03b4\u03cc\u03b8\u03b7\u03ba\u03b5 \u03b5\u03c0\u03b5\u03b9\u03b4\u03ae \u03bf\u03b9 \u0393\u03ac\u03bb\u03bb\u03bf\u03b9 \u03b1\u03c0\u03ad\u03ba\u03c1\u03c5\u03c0\u03c4\u03b1\u03bd \u03c4\u03b7\u03bd \u03bd\u03cc\u03c3\u03bf.<\/em><\/p>\n

\u0392\u03b9\u03b2\u03bb\u03b9\u03bf\u03b3\u03c1\u03b1\u03c6\u03af\u03b1<\/p>\n

Huai-Dong Song, Chang-Chun Tu, Guo-Wei Zhang, Sheng-Yue Wang, Kui Zheng, Lian-Cheng Lei, Qiu-Xia Chen,\u2026 and Guo-Ping Zhao. (2005) Cross-host evolution of severe acute respiratory syndrome coronavirus in palm civet and human. PNAS 102 (7) 2430-2435; https:\/\/doi.org\/10.1073\/pnas.0409608102<\/a><\/p>\n

Susanna K. P. Lau, Patrick C., Kenneth S. M. Li, Yi Huang\u2026 and Kwok-Yung Yuen, (2005) Severe acute respiratory syndrome coronavirus-like virus in Chinese horseshoe bats. PNAS vol. 102 no. 39 , 14040\u201314045<\/p>\n

Meriadeg Ar Gouilh, S\u00e9bastien J. Puechmaille, Jean-Paul Gonzalez , Emma Teeling , Pattamaporn Kittayapong , Jean-Claude Manuguerra. (2011) SARS-Coronavirus ancestor\u2019s foot-prints in South-East Asian bat colonies and the refuge theory. Infection, Genetics and Evolution 11, 1690\u20131702<\/p>\n

Jan Felix Drexler, Victor Max Corman, Christian Drosten (2014) Ecology, evolution and classification of bat coronaviruses in the aftermath of SARS. Antiviral Research 101, 45\u201356<\/p>\n

Ben Hu, Xingyi Ge, Lin-Fa Wang and Zhengli Shi. (2015) Bat origin of human coronaviruses, Virology Journal 12:221<\/p>\n

Vineet D. Menachery, Boyd L. Yount Jr., Amy C. Sims, Kari Debbink,, Sudhakar S. Agnihothram, \u2026, and Ralph S. Baric, (2016) SARS-like WIV1-CoV poised for human emergence. PNAS,vol. 113. 11,3048\u20133053, www.pnas.org\/cgi\/doi\/10.1073\/pnas.1517719113<\/a><\/p>\n

Jie\u00a0Cui , Fang\u00a0Li & Zheng- Li\u00a0Shi (2019) Origin and evolution of pathogenic coronaviruses. Nature Reviews Microbiology \u00a0vol 17.181<\/p>\n

WestonS, Frieman MB. (2020) COVID19: knowns, unknowns and questions. mSphere5 :e00203-20. https:\/\/doi.org\/10.1128\/ mSphere.00203-20.<\/p>\n

\n\t\t\t\t\t\n\t\t\t\t\t\t\"Fb-Button\"\n\t\t\t\t\t<\/a>\n\t\t\t\t<\/div>
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<\/div><\/div>","protected":false},"excerpt":{"rendered":"

\u0397 \u03bd\u03ad\u03b1 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b1 \u03b5\u03af\u03c7\u03b5 \u03c0\u03c1\u03bf\u03b2\u03bb\u03b5\u03c6\u03b8\u03b5\u03af. \u039f Menachery \u03ba\u03b1\u03b9 \u03bf\u03b9 \u03c3\u03c5\u03bd\u03b5\u03c1\u03b3\u03ac\u03c4\u03b5\u03c2 \u03c4\u03bf\u03c5, \u03c0.\u03c7. \u03b1\u03c0\u03cc \u03c4\u03bf 2016, \u03bc\u03b5 \u03c0\u03b5\u03b9\u03c1\u03ac\u03bc\u03b1\u03c4\u03b1 \u03b3\u03b5\u03bd\u03b5\u03c4\u03b9\u03ba\u03ae\u03c2 \u03bc\u03b7\u03c7\u03b1\u03bd\u03b9\u03ba\u03ae\u03c2 \u03b5\u03af\u03c7\u03b1\u03bd \u03c0\u03c1\u03bf\u03b2\u03bb\u03ad\u03c8\u03b5\u03b9 \u03cc\u03c4\u03b9 \u03ba\u03ac\u03c0\u03bf\u03b9\u03bf\u03b9 \u03ba\u03bf\u03c1\u03bf\u03bd\u03b1\u03ca\u03bf\u03af \u03c4\u03c9\u03bd \u03bd\u03c5\u03c7\u03c4\u03b5\u03c1\u03af\u03b4\u03c9\u03bd \u03b1\u03c0\u03cc \u03c4\u03b7\u03bd \u0393\u03b9\u03bf\u03c5\u03c7\u03ac\u03bd \u03bc\u03c0\u03bf\u03c1\u03b5\u03af \u03bd\u03b1 \u03c0\u03c1\u03bf\u03ba\u03b1\u03bb\u03ad\u03c3\u03bf\u03c5\u03bd \u03bd\u03ad\u03b5\u03c2 \u03b5\u03c0\u03b9\u03b4\u03b7\u03bc\u03af\u03b5\u03c2 \u03ba\u03b1\u03b9 \u03bc\u03b5\u03b3\u03ac\u03bb\u03b5\u03c2 \u03ba\u03c1\u03af\u03c3\u03b5\u03b9\u03c2. <\/p>\n

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